Ota, R. P., Lima, F. C. T. and Pavanelli, C. S. (2015) “A new species of Hemigrammus Gill, 1858 (Characiformes: Characidae) from the central and western Amazon and rio Paraná-Paraguai basins”, Zootaxa. Auckland, New Zealand, 3948(2), pp. 218–232. doi: 10.11646/zootaxa.3948.2.4.