Crawford, Andrew J., Roberto Alonso, César A. Jaramillo A., Samuel Sucre, and Roberto Ibáñez D. 2011. “DNA Barcoding Identifies a Third Invasive Species of Eleutherodactylus (Anura: Eleutherodactylidae) in Panama City, Panama”. Zootaxa 2890 (1). Auckland, New Zealand:65–67. https://doi.org/10.11646/zootaxa.2890.1.6.