Shapovalov, Andrey M., Oliver Hillert, David Král, Sergey V. Kolov, Mikhail F. Bagaturov, and Georgy V. Nikolajev. 2026. “Revision of the Lethrus (Heteroplistodus) crenulatus-Species Group (Coleoptera: Geotrupidae: Lethrinae). Part 1: Species Similar to L. tschitscherini Semenov, 1894”. Zootaxa 5831 (4). Auckland, New Zealand:451-76. https://doi.org/10.11646/zootaxa.5831.4.1.