Heinicke, Matthew P., César L. Barrio-Amorós, and S. Blair Hedges. 2015. “Molecular and Morphological Data Support Recognition of a New Genus of New World Direct-Developing Frog (Anura: Terrarana) from an under-Sampled Region of South America”. Zootaxa 3986 (2). Auckland, New Zealand:151–172. https://doi.org/10.11646/zootaxa.3986.2.1.