Dehling, J. Maximilian, and Masafumi Matsui. 2013. “A New Species of Leptolalax (Anura: Megophryidae) from Gunung Mulu National Park, Sarawak, East Malaysia (Borneo)”. Zootaxa 3670 (1). Auckland, New Zealand:33–44. https://doi.org/10.11646/zootaxa.3670.1.2.