Guayasamin, Juan M., and Alejandro F. Arteaga. 2013. “A New Species of the Pristimantis orestes Group (Amphibia: Strabomantidae) from the High Andes of Ecuador, Reserva Mazar”. Zootaxa 3616 (4). Auckland, New Zealand:345–356. https://doi.org/10.11646/zootaxa.3616.4.3.