MADDISON, D. R.; ARNOLD, A. E. A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species. Zootaxa, Auckland, New Zealand, v. 2214, n. 1, p. 45–61, 2009. DOI: 10.11646/zootaxa.2214.1.3. Disponível em: https://www.mapress.com/zt/article/view/zootaxa.2214.1.3. Acesso em: 6 sep. 2025.