Rödel, M.-O., Schmitz, A., Penner, J., Griesbaum, F., Leaché, A. D., Ofori-Boateng, C., Portik, D., Doumbia, J., Kouamé, N. G., Kpan, T. F., Gonwouo, N. L., Nopper, J., Hillers, A., Nago, S. G. A., Ernst, R., Glos, J., Petzold, A., & Barej, M. F. (2026). Four new cryptic Ptychadena species from the Upper Guinean rainforests of West Africa (Amphibia, Anura, Ptychadenidae). Zootaxa, 5779(1), 1–66. https://doi.org/10.11646/zootaxa.5779.1.1