Hutter, C. R., Lambert, S. M., Cobb, K. A., Andriampenomanana, Z. F., & Vences, M. (2015).

A new species of bright-eyed treefrog (Mantellidae) from Madagascar, with comments on call evolution and patterns of syntopy in the Boophis ankaratra complex

. Zootaxa, 4034(3), 531–555. https://doi.org/10.11646/zootaxa.4034.3.6