Schawaller, W. (2013). Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue . Zootaxa, 3721(4), 351–364. https://doi.org/10.11646/zootaxa.3721.4.3