Deschodt, C. M., & Davis, A. L. (2019).

New southern African species and a revalidation in the dung beetle genus Gyronotus van Lansberge, 1874 (Scarabaeidae: Scarabaeinae) with an updated key

. Zootaxa, 4624(2), 275–282. https://doi.org/10.11646/zootaxa.4624.2.10