Kohlmann, B., & Solís, Ángel. (2012).

New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama

. Zootaxa, 3193(1), 28–52. https://doi.org/10.11646/zootaxa.3193.1.2