Chaves-Acuña, W.; Chaves, G.; Klank, J.; Arias, E.; Bolaños, F.; Shepack, A.; Leenders, T.; Cossel, J.; Faivovich, J. Recent Findings of Isthmohyla Pictipes (Anura: Hylidae) in Costa Rica: Variation and Implications for Conservation. Zootaxa 2020, 4881, 499–514.