Celante, G.L., Carvalho, A.G., Jahyny, B.J.B., Araújo, K.B.S. and Santos, P.B.D. 2024. A new species of Athyreus MacLeay, 1819 (Coleoptera: Geotrupidae: Bolboceratinae: Athyreini) from northeastern Brazil. Zootaxa. 5512, 3 (Sep. 2024), 389–401. DOI:https://doi.org/10.11646/zootaxa.5512.3.4.