Jacobsen, I.P. and Bennett, M.B. 2009. A Taxonomic Review of the Australian Butterfly Ray Gymnura australis (Ramsay & Ogilby, 1886) and Other Members of the family Gymnuridae (Order Rajiformes) from the Indo-West Pacific. Zootaxa. 2228, 1 (Sep. 2009), 1–28. DOI:https://doi.org/10.11646/zootaxa.2228.1.1.