de Padua Almeida, A. and Angulo, A. 2006. A new species of Leptodactylus (Anura: Leptodactylidae) from the state of Espirito Santo, Brazil, with remarks on the systematics of associated populations. Zootaxa. 1334, 1 (Oct. 2006), 1–25. DOI:https://doi.org/10.11646/zootaxa.1334.1.1.