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Issue: Vol. 5869 No. 2: 17 Aug. 2026
Type: Article
Published: 2026-08-17
DOI: 10.11646/zootaxa.5869.2.1
Page range: 201-239
Abstract views: 384
PDF downloaded: 212

New species and records of Cirratulidae (Annelida, Polychaeta) from methane seep and seamount habitats in the Pacific Ocean off Costa Rica

Aquatic Research & Consulting; 24 Hitty Tom Road; Duxbury; MA 02332 USA; Department of Invertebrate Zoology; Museum of Comparative Zoology; Harvard University; 26 Oxford Street; Cambridge; MA 02138 USA
Scripps Institution of Oceanography; University of California San Diego; La Jolla; CA 92093-0202 USA
Scripps Institution of Oceanography; University of California San Diego; La Jolla; CA 92093-0202 USA
Annelida Aphelochaeta Chaetocirratulus Cirratulus Kirkegaardia Raricirrus deep-sea

Abstract

Eight species of polychaetes in six genera belonging to the family Cirratulidae are reported from seamounts and methane seeps off Costa Rica in the Pacific Ocean. Seven species are new to science. The genus Aphelochaeta includes two new species: A. pigmentata sp. nov., and A. rotunda sp. nov. The genera Chaetocirratulus, Chaetozone, Kirkegaardia, and Raricirrus each include a single new species: Chaetocirratulus chromatus sp. nov., Chaetozone carinata sp. nov., K. davidgeorgei sp. nov., and R. hartmanae sp. nov. The genus Cirratulus includes two species: C. megalus Chamberlin, 1919 originally described from deep water off Peru, and Cirratulus jacoensis sp. nov. from Jacó Summit off Costa Rica. The samples in which these taxa were found are all from deep water (>200 m) with most from bathyal depths of 1000–2184 m. Cirratulus tumbesiensis Carrasco, 1978 from Chile is referred to the genus Cirriformia.

References

  1. Augener, H. (1932) Polychaeten von den Auckland und Campbell-Inseln. Videnskabelige Meddelelser Naturhistorisk Forening i København, 75, 1–115, 44 figs. [https://biodiversitylibrary.org/page/27905256]
  2. Banse, K. & Hobson, K. (1968) Benthic polychaetes from Puget Sound, Washington. Proceedings of the United States National Museum, 125 (3667), 1–53. https://doi.org/10.5479/si.00963801.125-3667.1
  3. Berkeley, E. (1929) Polychaetous annelids from the Nanaimo District. 4. Chaetopteridae to Maldanidae. Contributions to Canadian Biology, Ottawa, new series, 4, 305–316, 1 plate. https://doi.org/10.1139/f29-022
  4. Berkeley, E. & Berkeley, C. (1941) On a collection of Polychaeta from Southern California. Bulletin of the Southern California Academy of Sciences, 40, 16–60. [https://www.biodiversitylibrary.org/page/34209137]
  5. Berkeley, E. & Berkeley, C. (1954) Notes on the life-history of the polychaete Dodecaceria fewkesi (nom. n.). Journal of the Fisheries Research Board of Canada, 11, 326–334. https://doi.org/10.1139/f54-021
  6. Blake, J.A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M.E. & Kirkegaard, J.B. (Eds.), Systematics, Biology and Morphology of World Polychaeta. Proceedings of the 2nd International Polychaete Conference, Copenhagen 1986. Ophelia, 5 (Supplement), 17–30. https://doi.org/10.1163/9789004629745_005
  7. Blake, J.A. (1996) Chapter 8. Family Cirratulidae Ryckholdt, 1851. Including a revision of the genera and species from the eastern North Pacific. In: Blake, J.A., Hilbig, B. & Scott, P.H. (Eds.), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and the Western Santa Barbara Channel. Volume 6. The Annelida Part 3, Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, Santa Barbara, California, pp. 263–384.
  8. Blake, J.A. (2006) New species and records of deep-water Cirratulidae (Polychaeta) from off Northern California. Scientia Marina, 70S3, 45–57. https://doi.org/10.3989/scimar.2006.70s345
  9. Blake, J.A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501–552. https://doi.org/10.11646/zootaxa.3919.3.5
  10. Blake, J.A. (2016) Kirkegaardia (Polychaeta, Cirratulidae), new name for Monticellina Laubier, preoccupied in the Rhabdocoela, together with new records and descriptions of eight previously known and sixteen new species from the Atlantic, Pacific, and Southern Oceans. Zootaxa, 4166 (1), 1–93. https://doi.org/10.11646/zootaxa.4166.1.1
  11. Blake, J.A. (2018) Bitentaculate Cirratulidae (Annelida, Polychaeta) collected chiefly during cruises of the R/V Anton Bruun, USNS Eltanin, USCG Glacier, R/V Hero, RVIB Nathaniel B. Palmer, and R/V Polarstern from the Southern Ocean, Antarctica, and off Western South America. Zootaxa, 4537 (1), 1–130. https://doi.org/10.11646/zootaxa.4537.1.1
  12. Blake, J.A. (2019) New species of Cirratulidae (Annelida, Polychaeta) from abyssal depths of the Clarion-Clipperton Fracture Zone, North Equatorial Pacific Ocean. Zootaxa, 4629 (2), 151–187. https://doi.org/10.11646/zootaxa.4629.2.1
  13. Blake, J.A. (2022) New species and records of Caulleriella, Chaetocirratulus, and Chaetozone (Annelida, Cirratulidae) from continental shelf and slope depths of the Western North Atlantic Ocean. Zootaxa, 5113 (1), 1–89. https://doi.org/10.11646/zootaxa.5113.1.1
  14. Blake, J.A. (2023) New species of Cirratulidae (Annelida) from Continental slope and abyssal depths off Eastern Australia. Records of the Australian Museum, 75, 249–270. https://doi.org/10.3853/j.2201-4349.75.2023.1799
  15. Blake, J.A. & Magalhães, W. (2019) Cirratulidae Ryckholt, 1851. In: Purschke, G., Böggemann, M. & Westheide, W. (Eds.), Handbook of Zoology. Annelida 1: Annelida Basal groups and Pleistoannelida, Sedentaria 1. pp. 339–397. https://doi.org/10.1515/9783110291582
  16. Buzhinskaja, G.N. & Smirnov, R.V. (2017) A new species of Raricirrus (Polychaeta, Ctenodrilidae) from the continental slope of the Laptev Sea near the Gakkel Ridge. Proceedings of the Zoological Institute RAS, 321 (4), 425–432. https://doi.org/10.31610/trudyzin/2017.321.4.425
  17. Carrasco, F.D. (1978) Polychaeta (Annelida) de Bahía de Concepción, Chile. Familias Orbiniidae, Cirratulidae, Cossuridae, Capitellidae y Ampharetidae, con la descripcion de tres especies y una subespecie nuevas. Boletín de la Sociedad de Biología de Concepción, 51 (1), 67–92. [https://bolsocbiolconcepc.cl/pdfs/v51(1)/10-Carrasco.pdf]
  18. Chamberlin, R.V. (1918) Polychaetes from Monterey Bay. Proceedings of the Biological Society of Washington, 31, 173–180. [https://www.biodiversitylibrary.org/page/3332088]
  19. Chamberlin, R.V. (1919) The Annelida Polychaeta. Memoirs of the Museum of Comparative Zoology, Harvard, 48, 1–514, pls. 1–80. [https://www.biodiversitylibrary.org/ia/memoirsofmuseumo4801harv]
  20. Chevez-Lopez, Y. & Cruz-Gomez, C. (2019) New records of polychaetes (Annelida, Polychaeta) from three locations of Oaxaca, Mexico. Revista de Biología Tropical, 67 (S5), Supplement, 157–168. https://doi.org/10.15517/rbt.v67iS5.38941
  21. Çinar, M.E. & Petersen, M.E. (2011) Re-description of Cirratulus dollfusi (Polychaeta: Cirratulidae), and Fauvelicirratulus as a new genus. Journal of the Marine Biological Association of the United Kingdom, 91 (2), 415–418. https://doi.org/10.1017/S0025315410000895
  22. Claparède, A.R.É. (1863) Beobachtungen über Anatomie und Entwicklungsgeschichte wirbelloser Thiere an der Küste von Normandie angestellt. Verlag von Wilhelm Engelmann, Leipzig, 120 pp., 18 plates. https://doi.org/10.5962/bhl.title.10030
  23. Czerniavsky, V. (1881) Materialia ad zoographiam Ponticam comparatam. Fasc. III Vermes [Second part]. Bulletin de la Société Impériale des Naturalistes de Moscou, 56 (2), 338–420. [https://biodiversitylibrary.org/page/41340542]
  24. Dean, H.K. (1995) A new species of Raricirrus (Polychaeta: Ctenodrilidae) from wood collected in the Tongue of the Ocean, Virgin Islands. Proceedings of the Biological Society of Washington, 108 (2), 169–179. [https://www.biodiversitylibrary.org/page/34571999]
  25. Dean, H.K. & Blake, J.A. (2007) Chaetozone and Caulleriella (Polychaeta: Cirratulidae) from the Pacific Coast of Costa Rica, with description of eight new species. Zootaxa, 1451, 41–68. https://doi.org/10.11646/zootaxa.1451.1.2
  26. Dean, H.K. & Blake, J.A. (2009) Monticellina (Polychaeta: Cirratulidae) from the Pacific coast of Costa Rica with descriptions of six new species. Zoosymposia, 2, 105–126. https://doi.org/10.11646/zoosymposia.2.1.10
  27. Dean, H.K. & Blake, J.A. (2016) Aphelochaeta (Polychaeta: Cirratulidae) from the Pacific coast of Costa Rica, with a description of five new species. Zootaxa, 4103 (2), 101–116. https://doi.org/10.11646/zootaxa.4103.2.1
  28. Díaz-Díaz, O. & Salazar-Vallejo, S.I. (2009) Chapter 12. Cirratulidae Ryckholt, 1851. In: de León-González, J.A., Bastida-Zavala, J.R., Carrera-Para, L.F., García-Garza, M.E., Peña-Rivera, A., Salazar-Vallejo, S.I. & Solís-Weiss, V. (Eds.), Poliquetos (Annelida, Polychaeta) de México y América Tropical. Vol. 1. Universidad Autónoma de Nuevo Leon, Monterrey, México, pp. 131–147.
  29. Díaz-Díaz, O. & Salazar-Vallejo, S.I. (2021) Chapter 13. Cirratulidae Ryckholt, 1851. In: de León-González, J.A., Bastida-Zavala, J.R., Carrera-Para, L.F., García-Garza, M.E., Salazar-Vallejo, S.I., Solís-Weiss, V. & Tovar-Hernández, M.A. (Eds.), Anélidos Marinos de México y América Tropical. Universidad Autónoma de Nuevo Leon, Monterrey, México, pp. 187–207.
  30. Doner, S.A. & Blake, J.A. (2009) Two new species of Aphelochaeta (Polychaeta: Cirratulidae) from deep-water off northern California. Zoosymposia, 2, 127–137. https://doi.org/10.11646/zoosymposia.2.1.11
  31. Eilertsen, M.H., Songsrud, J.A., Tandberg, A.H.S., Alvestad, T., Budaeva, N.M. & et al. (2024) Diversity, habitat endemicity and trophic ecology of the fauna of Loki’s Castle vent field on the Arctic Mid-Ocean Ridge. Scientific Reports, 14, 103. https://doi.org/10.1038/s41598-023-46434-z
  32. Freitas, R., Ribeiro, R.P. & Ruta, C. (2022) Kirkegaardia Blake, 2016 (Annelida: Cirratulidae) from Southeastern Brazil with description of nine new species. Plos One, 17 (5), e0265336. https://doi.org/10.1371/journal.pone.0265336
  33. Fauchald, K. (1972) Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1–575, 68 plates. [https://repository.si.edu/handle/10088/6207]
  34. Grosse, M., Kongsrud, J.A., Alvestad, T., Eilertsen, M.H. & Budaeva, N. (2025) A new species and a new record of Cirratulidae (Annelida, Cirratulida) from Loki’s vent field. European Journal of Taxonomy, 987, 1–25. https://doi.org/10.5852/ejt.2025.987.2855
  35. Hartman, O. (1936) Nomenclatural changes involving California polychaete worms. Journal of the Washington Academy of Sciences, 26, 31–32. [https://biodiversitylibrary.org/page/39872214]
  36. Hartman, O. (1948) The polychaetous annelids of Alaska. Pacific Science, 2, 1–58. [http://hdl.handle.net/10125/8890]
  37. Hartman, O. (1960) Systematic account of some marine invertebrate animals from the deep basins of Southern California. Allan Hancock Pacific Expeditions, 22, 69–215. [https://www.biodiversitylibrary.org/page/4682240]
  38. Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 25, 1–226, 34 plates. [https://www.biodiversitylibrary.org/page/5214831]
  39. Hartman, O. (1963) Submarine Canyons of Southern California. Part III. Systematics: Polychaetes. Allan Hancock Pacific Expeditions, 27 (3), 1–93. [https://www.biodiversitylibrary.org/page/4683354]
  40. Hartman, O. (1969) Atlas of the Sedentariate Polychaetous Annelids from California. Allan Hancock Foundation, University of Southern California, Los Angeles, 812 pp.
  41. Hiley, A.S., Green, K.R. & Rouse, G.W. (2025) Seven new species of scaleworms (Lepidonotopodini, Polynoidae, Polychaeta, Annelida) from deep-sea chemosynthetic-based ecosystems. Marine Biodiversity, 55, 106. https://doi.org/10.1007/s12526-025-01580-7
  42. Jimi, N., Fujimoto, S., Yoshihiro, F., Oguchi, K. & Miura, T. (2022) Four new species of Ctenodrilus, Raphidrilus, and Raricirrus (Cirratuliformia, Annelida) in Japanese waters, with notes on the phylogenetic position. PeerJ, 10, e13044. https://doi.org/10.7717/peerj.13044
  43. Johnson, H.P. (1901) The Polychaeta of the Puget Sound Region. Proceedings of the Boston Society of Natural History, 29 (1), 381–437. [https://www.biodiversitylibrary.org/page/56803706]
  44. Kinberg, J.G.H. (1866) Annulata Nova. Öfversigt af Königlich Vetenskapsakademiens förhandlingar, Stockholm, 22 (4), 239–258. [https://biodiversitylibrary.org/page/32339515]
  45. Laubier, L. (1961) Monticellina heterochaeta n. g., sp. nov., Cténodrilide (Polychètes Sedéntaires) des vases côtieres de Banyuls sur-Mer. Vie et Milieu, 11, 601–604. [https://wwwphp.obs-banyuls.fr/Viemilieu/images/Archives_1950-2002/vie-et-milieu/VOLUME_1960_11_fasc4.pdf]
  46. Lamarck, J.B. de. (1818) Histoire naturelle des Animaux sans Vertèbres, préséntant les caractères généraux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales espèces qui s’y rapportent; precedes d’une Introduction offrant la determination des caracteres essentiels de l`Animal, sa distinction du vegetal et desautres corps naturels, enfin, l’Exposition des Principes fondamentaux de la Zoologie. Deterville, Paris, 612 pp. [http://biodiversitylibrary.org/page/12886879]
  47. Langerhans, P. (1881) Die Wurmfauna von Madeira. III. Zeitschrift für wissenschaftliche Zoologie, 34 (1), 87–143. [https://biodiversitylibrary.org/page/42353743]
  48. Magalhães, W.F. & Bailey-Brock, J.H. (2013) Bitentaculate Cirratulidae (Annelida: Polychaeta) from the northwestern Pacific Islands with a description of nine new species. Zootaxa, 3630 (1), 80–116. https://doi.org/10.11646/zootaxa.3630.1.3
  49. Magalhães, W.F., Linse, K. & Wiklund, H. (2017) A new species of Raricirrus (Annelida: Cirratuliformia) from deep-water sunken wood off California. Zootaxa, 4353 (1), 51–68. https://doi.org/10.11646/zootaxa.4353.1.3
  50. Malmgren, A.J. (1867) Annulata Polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Öfversigt af Kongliga Vetenskaps-Akademien Förhandlingar, 24, 127–235. https://doi.org/10.5962/bhl.title.13358
  51. Monticelli, F.S. (1910) Di un nuovo Ctenodrilide del Golfo di Napoli. Rendiconto dell’Accademia delle Scienze Fisiche e Matematiche (Sezione della Società Reale di Napoli), Serie 3, 16 (3–4), 61–64. [https://babel.hathitrust.org/cgi/pt?id=njp.32101064104308&seq=505]
  52. Montagne, D. (1984) Expanded definition of Raricirrus maculatus. SCAMIT Newsletter, 3 (6), 1–2. [https://scamit.org/newsletters/1984-09.pdf]
  53. Montagu, G. (1808) Description of several marine animals found on the south coast of Devonshire. Transactions of the Linnean Society of London, 9, 81–114, plates 2–8. https://doi.org/10.1111/j.1096-3642.1818.tb00327.x
  54. Moore, J.P. (1903) Polychaeta from the coastal slope of Japan and from Kamchatka and Bering Sea. Proceedings of the Philadelphia Academy of Natural Sciences, 55, 401–490, plates 23–27. [https://www.biodiversitylibrary.org/page/6600785]
  55. Moore, J.P. (1904) New Polychaeta from California. Proceedings of the Philadelphia Academy of Natural Sciences, 56, 484–503, plates 37–38. [https://biodiversitylibrary.org/page/6367345]
  56. Moore, J.P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the of the Philadelphia Academy of Natural Sciences, 61, 235–295, plates 7–9. [https://www.biodiversitylibrary.org/page/12715216]
  57. Moore, J.P. (1923) Polychaetous annelids dredged by the U.S.S. Albatross off the coast of southern California in 1904. Spionidae to Sabellariidae. Proceedings of the Philadelphia Academy of Natural Sciences, 75, 179–259, plates 17–18. https://doi.org/10.5962/bhl.title.12425
  58. Müller, O.F. (1776) Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Havniae, 282 pp. https://doi.org/10.5962/bhl.title.13268
  59. Örsted, A.S. (1843) Annulatorum danicorum conspectus. Naturhistorisk Tideskrift København, 4, 1–52, 8 text figs. + 7 plates. Maricolae. Copenhagen, 52 pp. https://doi.org/10.5962/bhl.title.11849
  60. Petersen, M.E. (1991) A review of asexual reproduction in Cirratulidae (Annelida: Polychaeta), with redescription of Cirratulus gayheadius (Hartman, 1965), new combination and emendation or reinstatement of some cirratulid genera. In: Reish, D.J. (Ed.), Third International Polychaete Conference held at California State University, Long Beach, California August 1969. Bulletin of Marine Science, 48 (2), 592 (Abstract).
  61. Petersen, M.E. & George, J.D. (1991) A new species of Raricirrus from Northern Europe, with notes on its biology and discussion of the affinities of the genus (Polychaeta: Ctenodrilidae). In: Petersen, M.E. & Kirkegaard, J.B. (Eds.), Systematics, Biology and Morphology of World Polychaeta. Proceedings of the 2nd International Polychaete Conference, Copenhagen 1986. Ophelia, Supplement 5, 185–208. https://doi.org/10.1163/9789004629745_019
  62. Read, G. & Fauchald, K. (Ed.) (2025a) World Polychaeta Database. Cirratulidae Ryckholt, 1851. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=919 (accessed 24 May 2025)
  63. Read, G. & Fauchald, K. (Ed.) (2025b) World Polychaeta Database. Aphelochaeta Blake, 1991. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=129240 (accessed 24 May 2025)
  64. Read, G. & Fauchald, K. (Ed.) (2025c) World Polychaeta Database. Chaetocirratulus Blake, 2018. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=1316610 (accessed 19 April 2025)
  65. Read, G. & Fauchald, K. (Ed.) (2025d) World Polychaeta Database. Kirkegaardia Blake, 2016. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=884676 (accessed 10 April 2025)
  66. Read, G. & Fauchald, K. (Ed.) (2025e) World Polychaeta Database. Cirratulus Lamarck, 1818. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=129243 (accessed 28 May 2025)
  67. Read, G. & Fauchald, K. (Ed.) (2025f) World Polychaeta Database. Raricirrus Hartman, 1961. World Register of Marine Species, Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=129254 (accessed 01 Mar 2025)
  68. Rioja, E. (1960) Estudios Anelidologicos. XXIII. Contribucion al conocimiento de Los Anelidos poliquetos de las Islas de Revillagigedo. Anales del Instituto de Biología, 37 (1–2), 243–259. [https://anales.ib.unam.mx/index.php/anales/article/view/1136/1282]
  69. Ryckholt, Philippe de (Baron) (1851) Mélanges paléontologiques. Part 1. Memoires Couronnes et Memoires des Savants Etrangers de l’Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 24, 1–176, pls. 1–10. [https://biodiversitylibrary.org/page/2718124]
  70. Seid, C.A., Hiley, A.S., McCowin, M.F., Carvajal, J.I., Cha, H., Ahyong, S.T., Ashford, O.S., Odalisca, B., Eernisse, D.J., Goffredi, S.K., Hendrickx, M.E., Kocot, M., Mah, L., Miller, A.K., Mongiardino, N., Mooi, R., O’Hara, T.D., Pleijel, F., Stiller, J., Tilic, E., Valentich-Scott, P., Warén, A., Wicksten, M.K., Wilson, N.G., Cordes, E.E., Levin, L.A., Cortés, J. & Rouse, G.W. (2025) A faunal inventory of methane seeps on the Pacific margin of Costa Rica. Zookeys, 1222, 1–250. https://doi.org/10.3897/zookeys.1222.134385
  71. Schmidt, O. (1857) Zur Kenntnis der Turbellaria rhabdocoela und einiger anderer Würmer des Mittlelmeeres. Sitzungsberichte der Kaiserliche Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Klasse, 23, 347–366, pls. 1–5. [http://biodiversitylibrary.org/page/36386704]
  72. Tasso, V., Haddad, M.E., Assadi, C., Canales, R., Aguirre, L. & Velez-Auaxo, X. (2018) Macrobenthic fauna from an upwelling coastal area of Peru (Warm Temperate South-eastern Pacific province–Humboldtian ecoregion). Biodiversity Data Journal, 6, e2893e7. https://doi.org/10.3897/BDJ.6.e2893e7
  73. Westblad, E. (1953) New Turbellaria parasites of echinoderms. Arkiv för Zoologi, Series 2, 5, 269–288.
  74. Webster, H.E. & Benedict, J.E. (1887) The Annelida Chaetopoda, from Eastport, Maine. U.S. Commission of Fish & Fisheries. Report of the United States Commissioner of Fisheries. 1885. Part 13, II. Appendix to report of Commissioner, D.22. 707–758, pls. 1–8. [http://biodiversitylibrary.org/page/15839855]
  75. Wilfert, M. (1974) Aphropharynx heterochaeta nov. gen. nov. spec., ein neuer polychaet aus der familie Ctenodrilidae Kennel 1882. Cahiers de Biologie Marine, 15 (4), 495–504. https://doi.org/10.21411/CBM.A.F5889AF8

How to Cite

Blake, J.A., Seid, C.A. & Rouse, G.W. (2026) New species and records of Cirratulidae (Annelida, Polychaeta) from methane seep and seamount habitats in the Pacific Ocean off Costa Rica. Zootaxa, 5869 (2), 201–239. https://doi.org/10.11646/zootaxa.5869.2.1