Palaeoentomology <p><strong>Palaeoentomology </strong>is the official journal of the <a href="">International Palaeoentomological Society</a>. It is an international peer-reviewed scientific journal, which publishes high quality, original research contributions as well as review papers. Papers are published in English and they cover a wide spectrum of topics in palaeoentomology, fossil terrestrial arthropods and amber research, i.e. systematic palaeontology, morphology, diversity, palaeogeography, palaeoecology, palaeobehavior, evolutionary and phylogenetic studies on fossil insects and terrestrial arthropods, biostratigraphy, taphonomy, and amber (deposits, inclusions, geochemistry, curation). Descriptions of new methods (analytical, instrumental or numerical) should be relevant to the broad scope of the journal.</p> en-US <span lang="EN-GB">Authors need to complete and return an </span><span lang="EN-GB"><a href="/phytotaxa/images/copyright.rtf">Assignment of Copyright</a> </span><span lang="EN-GB">form when a paper is accepted for publication. Authors from institutions that do not allow transfer of copyrights to publishers (e.g. government institutions such as USDA, CSIRO) should attach a copyright waiver or similar document.</span> (Diying Huang) (Journal support team) Fri, 24 Sep 2021 13:59:35 +1200 OJS 60 <p><strong>A new leafhopper (Hemiptera: Cicadellidae) from late Eocene Rovno amber described based on an adult and associated last-instar nymph from Perebrody (Ukraine)</strong></p> <p><em>Rovnoxestus rasnitsyni</em> <strong>gen. &amp; sp. nov</strong>. is described from Eocene Rovno amber based on an adult female and fifth-instar nymph collected at a recently discovered locality at Perebrody, Rovno Province, Ukraine. The new fossil taxon is tentatively placed in Aphrodinae and resembles <em>Xestocephalites</em> Dietrich &amp; Gonçalves from Eocene Baltic amber but has the hind femur macrosetal formula 2+2+1 and hind tarsomere I in both nymph and adult with an elongated inner preapical seta. This is the first species of Eocene leafhopper for which both the adult and nymph are described in detail.</p> CHRISTOPHER H. DIETRICH, DMITRY A. DMITRIEV, EVGENY E. PERKOVSKY Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>New species of <em>Kamopanorpa</em> Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)</strong></p> <p>The systematic position of Microptysmatidae and Protomeropidae is critically reviewed in light of the latest reassessment by Minet <em>et al</em>. The Microptysmatidae belongs to a stem-group of Trichoptera, or at least to stem-Amphiesmenoptera (depending on chosen methodology), rather than to its own order Permotrichoptera as suggested by Minet <em>et al</em>., and Protomeropidae are not Mecoptera. The double anal loop of <em>Kamopanorpa </em>Martynov, 1928 is not significantly different from that of more advanced amphiesmenopterans, while in <em>Microptysmella</em> Kukalová-Peck &amp; Willmann, 1990, anal veins show clear tendency to looping (not a “fairly different arrangement”). If Microptysmatidae are considered in the traditional way, the polarity of characters listed by Minet <em>et al</em>. as supporting monophyly of the order Permotrichoptera can be treated just the opposite: divided veins RS<sub>1</sub>, RS<sub>2</sub> and M<sub>1</sub> are symplesiomorphies with Protomeropidae, and simple M<sub>4</sub> is the synapomorphy of Microptysmatidae, Cladochoristidae, and Amphiesmenoptera <em>s</em>.<em> str</em>. In addition, three new species of <em>Kamopanorpa </em>are described from the Permian deposits of Minusinsk Coal Basin, South Siberia: <em>Kamopanorpa rasnitsyni</em> <strong>sp. nov.</strong>, <em>K. sivchikovi</em> <strong>sp. nov.</strong>, and <em>K. rotunda</em> <strong>sp. nov.</strong> Some traits of fore- and hindwing venation of Microptysmatidae are discussed.</p> ALEXEY S. BASHKUEV, IRINA D. SUKATSHEVA Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A new species of the ʻprotozygopteranʼ damselfly (Odonata: Permagrionidae) from the Lower-Middle Permian of Russia</strong></p> <p>The small Paleozoic protozygopteran family Permagrionidae comprises 11 described species in 5 genera from the Lower Permian Chekarda and Solikamsk localities in Russia (Zalessky, 1948; Nel <em>et al</em>., 2012) and Salagou Formation in France (Nel <em>et al</em>., 1999; Fate <em>et al</em>., 2013), the Middle Permian Soyana and Kargala localities in Russia (Martynov, 1932; Martynov, 1937; Nel <em>et al</em>., 2012), and the Upper Permian Bodie Creek Head locality in Malvinas Tillyard (1928). Here we describe the new species, <em>Epilestes rasnitsyni </em><strong>sp. nov. </strong>from the Ufimian of Perm Territory, which is characterized by specific arrangement of veins in the petiole and the unique preservation of body structures.</p> ANASTASIA FELKER, DMITRY VASILENKO Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A Gondwanan record of the extinct genus <em>Cretobibio</em> (Diptera: Bibionidae)</strong></p> <p>A new species of <em>Cretobibio</em> Skartveit &amp; Ansorge, 2020, a genus formerly known only from the Lower Cretaceous of Spain, is described from the Lower Cretaceous of Brazil (Crato Formation, upper Aptian). A critical overview of scarce Mesozoic record of the Bibionidae is given.</p> ELENA DMITRIEVNA LUKASHEVICH, DALTON DE SOUZA AMORIM, GUILHERME CUNHA RIBEIRO Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Long-headed predators in Cretaceous amber—fossil findings of an unusual type of lacewing larva</strong></p> <p>Lacewing larvae (Neuroptera) are known to be fierce predators which are morphologically highly specialised for a raptorial lifestyle. Mandibular-maxillary stylets are characteristic for all larvae of this group; these stylets can be extraordinarily massive. Despite these distinct sucking-piercing stylets, also other extreme features occur in some ingroups, such as an extremely elongated neck. In larvae of thread-winged lacewings (Crocinae) the neck can reach up to about one third of the body length; they are also called ‘long-necked antlions’. Even though the larvae of living neuropteran species show a variety of conspicuous morphologies today, indeed 100 million years ago, in the Cretaceous, Neuroptera seems to have had an even more “experimental phase”. Several larval specimens are known so far especially in Myanmar, Spanish and Lebanese amber from the Cretaceous with unique and unusual character combinations not found in any group living today. We describe here ten new fossil findings of one of these types of larvae with elongated head capsule in Myanmar amber, previously only known from a single specimen. We compared the head shapes of the new specimens with those of 190 specimens of other lacewing larvae and discuss further implications of our findings, especially making functional comparisons with long-necked antlions.</p> ANA ZIPPEL, CHRISTINE KIESMÜLLER, GIDEON T. HAUG, PATRICK MÜLLER, THOMAS WEITERSCHAN, CAROLIN HAUG, MARIE K. HÖRNIG, JOACHIM T. HAUG Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Redescriptions of beetles of the <em>Notocupes</em> generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia</strong></p> <p>The term “generic complex” is proposed for 12 previously synonymized fossil genera of Ommatidae. Redescriptions of three species of the genus <em>Notocupes </em>are provided: <em>N</em>.<em> excellens</em> Ponomarenko, 1966, <em>N</em>.<em> caudatus</em> Ponomarenko, 1966 and <em>N</em>.<em> vitimensis</em> Ponomarenko, 1966, as well as a description of an elytron of a new species <em>Zygadenia alexrasnitsyni</em> <strong>sp. nov.</strong>, from the Lower Cretaceous locality Baissa (Vitim River headwaters, Eastern Transbaikalia). The descriptions contain selected morphological characters that will be used in future redescriptions.</p> OLESYA D. STRELNIKOVA, EVGENY V. YAN Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Discovery of a new genus and species of Dryinidae (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber: <em>Rasnitsynum burmense</em> gen. et sp. nov.</strong></p> <p>Dryinidae (Hymenoptera: Chrysidoidea) are parasitoids, and often also predators, of leafhoppers, planthoppers and treehoppers (Hemiptera: Auchenorrhyncha) (Guglielmino <em>et al</em>., 2013; Olmi <em>et al</em>., 2020b). Fossils of Dryinidae are known both as inclusions in amber and as compression fossils (Perkovsky <em>et al</em>., 2019; Olmi <em>et al</em>., 2010, 2020b; Martynova <em>et al</em>., 2020). Among the amber deposits, those situated in Myanmar are the richest. Mid-Cretaceous Burmese amber from Kachin State is known in fact to include Dryinidae belonging to the following subfamilies and genera: Anteoninae: <em>Burmanteon</em> Engel, 2003 (one species); Burmadryininae: <em>Burmadryinus</em> Olmi, Xu &amp; Guglielmino, 2014 (one species); Dryininae: <em>Dryinus</em> Latreille, 1804 (12 species) (Martynova <em>et al</em>., 2020); <em>Hybristodryinus</em> Engel, 2005 (17 species) (Perkovsky <em>et al</em>., 2019; Tribull <em>et al</em>., 2020; Olmi <em>et al</em>., 2021; Wang <em>et al</em>., 2021); <em>Pseudodryinus</em> Olmi, 1991 (one species); Palaeoanteoninae: <em>Palaeoanteon</em> Olmi, 2000 (one species) (Perkovsky <em>et al</em>., 2020a); Raptodryininae: <em>Raptodryinus</em> Olmi, Perkovsky, Martynova, Contarini, Bückle, Guglielmino, 2020 (one species).</p> MASSIMO OLMI, ADALGISA GUGLIELMINO, GIANLUCA PARISE, LEONARDO CAPRADOSSI, EVGENY E. PERKOVSKY Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A new species of the fossil genus <em>Electrotrichia</em> (Insecta: Trichoptera: Hydroptilidae) from Rovno amber (Zhytomyr region, Olevsk locality)</strong></p> <p>Recent discoveries of amber deposits have resulted in numerous new caddisfly species of Rovno amber described in a series of recent papers (Melnitsky &amp; Ivanov, 2010, 2013, 2016a, b). The list of species previously known from Rovno amber included 42 species (Ivanov <em>et al</em>., 2016; Perkovsky, 2017). Our study is based on a collection of Trichoptera from a new Rovno amber locality near Olevsk in Zhytomyr region. Nine amber species have been reported from this region (Legalov <em>et al</em>., 2021; Radchenko <em>et al</em>., 2021), including new ant species, a new anthribid, two new genera, and four new species of gall midges (Fedotova &amp; Perkovsky, 2015, 2017); additional new cicadellid genus and species is described from closely connected fauna of Perebrody (Dietrich <em>et al</em>., 2021). It total, the new collection includes 6 pieces of Rovno Amber with six specimens of Trichoptera, two of those are described below as a new species of <em>Electrotrichia</em>.</p> STANISLAV I. MELNITSKY, VLADIMIR D. IVANOV, EVGENY E. PERKOVSKY Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Re-description of the Cretaceous wasp <em>Humiryssus leucus</em> Lin, 1980 (Hymenoptera: Evanioidea: Baissidae)</strong></p> <p><em>Humiryssus</em> Lin, 1980 is a Cretaceous genus with <em>Humiryssus leucus</em> Lin, 1980 as its type species established based on a tiny wasp from the Lower Cretaceous (Hauterivian-Barremian) Laocun Formation at Laocun, Jiande, Zhejiang Province, eastern China and originally placed in the extinct family Paroryssidae (Hymenoptera: Tenthredinoidea) (Lin, 1980). It was later treated jointly with the genus <em>Manlaya</em> Rasnitsyn, 1980 (Evanioidea: Gasteruptiidae: Baissinae) because the characters shown in the line drawing by Lin (1980) indicate a close relationship of the genus to <em>Manlaya</em> (Rasnitsyn <em>et al</em>., 1998). However, Zhang &amp; Rasnitsyn (2004) considered it to be a separate genus within the Baissinae, which is now considered as an independent family under the Evanioidea (<em>e</em>.<em>g</em>., Li <em>et al</em>., 2018; Jouault <em>et al</em>., 2020, 2021).</p> HAICHUN ZHANG Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A new mayfly species <em>Triassodotes rasnitsyni</em> sp. nov. of the family Misthodotidae Tillyard, 1932 (Insecta; Ephemerida, Permoplectoptera) from the Triassic deposits of Kuzbass, Russia</strong></p> <p>The mayfly sister group family Misthodotidae Tillyard, 1932 includes two genera: <em>Misthodotes</em> Sellards, 1909 and <em>Triassodotes</em> Sinitshenkova &amp; Papier, 2005. <em>Misthodotes</em> species have so far been known only from the Permian, six Early Permian species have been described from North America (Carpenter, 1933, 1979; Tillyard, 1932, 1936), one from Germany (Kinzelbach &amp; Lutz, 1984) and two from the Perm region of Russia (Tshernova, 1965). Only three species are known from the Upper Permian deposits: two from the famous Isady locality in the Vologda Region of Russia (Sinitshenkova, 2013; Sinitshenkova &amp; Vassilenko, 2012) and one from the Urals (Novokshonov <em>et al</em>., 2002). The only <em>Triassodotes</em> species was found in the Middle Triassic Grès à Voltzia deposits in France (Sinitshenkova <em>et al</em>., 2005).</p> NINA D. SINITSHENKOVA Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A new genus of the family Alexarasniidae (Insecta: Embiida) from the Triassic of Kyrgyzstan</strong></p> <p>The Recent webspinners (Embiida = Embioptera) comprise a small order of insects; there are about 400 species distributed mainly in tropical and subtropical regions (Miller <em>et al</em>., 2012). They are gregarious, living subsocially in galleries of fine silk which they spin from glands on their forelegs (Edgerly <em>et al</em>., 2002). The webspinner lineage is traceable back via Middle Cretaceous Sorellembiidae, Jurassic Sinembiidae and Late Permian–Triassic Alexarasniidae to the Early–Middle Permian Atactophlebiidae (Engel &amp; Grimaldi, 2006; Huang &amp; Nel, 2009; Shcherbakov, 2015).</p> DANIIL S. ARISTOV, SERGEY YU. STOROZHENKO Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong><em>Aleksiana rasnitsyni</em> gen. et sp. nov. (Diptera, Limoniidae) to honour Alexandr Rasnitsyn</strong></p> <p>The family Limoniidae is the most speciose family of the infraorder Tipulomorpha, as well as one of the largest families of nematoceran Diptera. The oldest known representative of Limoniidae is <em>Architipula youngi</em> Krzemiński, 1992 described from the Upper Triassic of North America (<em>ca</em>. 208 Ma) belonging to the subfamily Architipulinae (Krzemiński, 1992). The subfamily Limoniinae (Limoniidae) stratigraphic range extends from the Lower Cretaceous Lebanese amber (Kania <em>et al</em>., 2014) to the present day, and is divided into two tribes, namely Antochini and Limoniini Savchenko (1985). Antochini currently comprise of the following contemporary genera: <em>Antocha</em> Alexander, 1924 (represented by 160 species); <em>Elliptera</em> Schiner, 1863 (represented by 12 species); <em>Orimarga</em> Osten-Sacken, 1869 (represented by 150 species) and <em>Thaumastoptera</em> Mik, 1866 (represented by 11 species). Representatives of this tribe currently occur on all continents except Antarctica, but individual genera are not distributed uniformly throughout the world (Oosterbroek, 2021). Although the family Limoniidae has been known since the Upper Triassic (Krzemińska &amp; Krzemiński, 2003), the oldest representative of the tribe Antochini is only known from the beginning of the Upper Cretaceous from the Burmese amber from Kachin (Podenas &amp; Poinar, 2009) dated to <em>ca</em>. 99 Ma (Shi <em>et al</em>., 2012).</p> WIESŁAW KRZEMIŃSKI, KATARZYNA KOPEĆ, AGNIESZKA SOSZYŃSKA-MAJ, KORNELIA SKIBIŃSKA Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>A new biting midge of the genus <em>Forcipomyia</em> Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae)</strong></p> <p>Biting midges (Ceratopogonidae) are a relatively well-studied family of nematocerous flies distributed worldwide, including 6,206 extant and 296 fossil species. To date, 1,146 extant and 32 fossil species in the genus <em>Forcipomyia</em> have been recorded in the world (Borkent &amp; Dominiak, 2020). Biting midges of the subgenus <em>Forcipomyia</em> s. str. are indicative of forests because their larvae and pupae usually live under the bark of rotting trees (Szadziewski, 2018). The oldest records of the genus are from the Eocene. <em>Forcipomyia</em> are reported from early Eocene Fushun amber (one species; Hong, 1981; Stebner <em>et al</em>., 2016; Szadziewski, 2018; Krzemiński <em>et al</em>., 2019), early Eocene Cambay amber (unnamed, Stebner <em>et al</em>., 2017), middle Eocene Sakhalin amber (one species, Szadziewski &amp; Sontag, 2013), Eocene Baltic amber from the Gulf of Gdańsk, Rovno and Bitterfeld (21 species; Szadziewski, 1988, Szadziewski, 1993, Sontag &amp; Szadziewski, 2011), Miocene Dominican amber (10 species, Szadziewski &amp; Grogan, 1998) and Miocene Mexican amber (unnamed, Szadziewski &amp; Grogan, 1996).</p> RYSZARD SZADZIEWSKI, ELŻBIETA SONTAG, MADELINE V. PANKOWSKI Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Special issue of “The 85<sup>th</sup> anniversary of Professor Alexandr P. Rasnitsyn” Part I (Front matter)</strong></p> DMITRY KOPYLOV Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>On the 85<sup>th</sup> anniversary of Alexandr Rasnitsyn</strong></p> <p>This issue of Palaeoentomology is dedicated to the foremost Russian entomologist, palaeontologist, and evolutionary biologist, Alexandr Pavlovich Rasnitsyn, who will be 85 on September 24 this year. The authors and those numerous colleagues, who could not, for various reasons, participate, wish a happy birthday to the undisputed worldwide leader of palaeoentomology!</p> ELENA D. LUKASHEVICH, ALEXEY S. BASHKUEV, BIDDY JARZEMBOWSKI, EDMUND A. JARZEMBOWSKI, ROMAN A. RAKITOV, DMITRY V. VASILENKO Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200 <p><strong>Alik Rasnitsyn, nearly 70 years together and apart</strong></p> <p>Of course, I have known for a long time that one has to call him by his full name, Alexandr Pavlovich, yet for me and a great many other people he remains Alik—despite the fact that I have never heard of anyone having ever questioned his scientific or moral authority. The following text is not a portrait or even a biography of A.P. Rasnitsyn, but merely what I have managed to put together from my recollections in defiance of my dementia. I am stuck in the country and cannot check basic facts about Alik, so, if I have forgotten or mixed up something, please forgive me.</p> ALEXANDR G. PONOMARENKO Copyright (c) 2021 Magnolia press limited Fri, 24 Sep 2021 00:00:00 +1200