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Issue: Vol. 21 No. 1: 7 Aug. 2026
Type: Article
Published: 2026-07-15
DOI: 10.11646/megataxa.21.1.1
Page range: 1-148
Abstract views: 729
PDF downloaded: 565

All genera of the world: order Opiliones (Animalia: Arthropoda: Arachnida)

Departamento de Invertebrados; Museu Nacional/Universidade Federal do Rio de Janeiro; Quinta da Boa Vista; São Cristóvão; 20.940-040 Rio de Janeiro (RJ); Brazil
Museum of Comparative Zoology & Department of Organismic and Evolutionary Biology; Harvard University; Cambridge; Massachusetts 02138; USA
Graduate Program in Geosciences; Rio de Janeiro State University; Rua São Francisco Xavier; 524; Maracanã; 20950-000; Rio de Janeiro; RJ; Brazil; Laboratório de Radioecologia e Mudanças Globais (LARAMG); Rio de Janeiro State University (UERJ)
Departamento de Vertebrados; Museu Nacional/Universidade Federal do Rio de Janeiro; Quinta da Boa Vista; São Cristóvão; 20.940-040 Rio de Janeiro (RJ); Brazil
Opiliones Laniatores Eupnoi Dyspnoi Cyphophthalmi nomenclature generic names synonyms type species harvestmen

Abstract

Opiliones is one of the most diverse orders of Chelicerata, ranking behind Araneae, Acariformes, and Parasitiformes in species richness. The group currently comprises approximately 6,900 valid and 1,500 invalid species, organized in about 1,700 valid and 800 invalid genera, yielding a remarkably low species-to-genus ratio of roughly 4:1 that reflects a history of naming monotypic genera based on a lack of phylogenetic understanding. In this contribution, we present an updated list of all genera of Opiliones, valid and invalid, organized by family-group taxa within each of the four suborders, Cyphophthalmi, Eupnoi, Dyspnoi and Lanitores. We also propose generic reallocations and nomenclatural corrections to improve the consistency of suprageneric classification. Specifically: the fossil genus † Halitherses Giribet & Dunlop, 2005 is placed in the new superfamily † Halithersoidea stat. nov. within Dyspnoi; the family Trionyxellidae Roewer, 1912 is synonymized with Assamiidae Sørensen, 1884; Goniosomoides Mello-Leitão, 1932 is synonymized with Deltaspidium Roewer, 1927, and Deltaspidium viridans (Mello-Leitão, 1932) comb. nov. is treated as a nomen inquirendum; the East Asian harvestman genus Pseudoliobunum A. Müller, 1914 is resurrected from the synonymy of Leiobunum C.L. Koch, 1839 under its correct original spelling, correcting the unjustified emendation Pseudoleiobunum Tan, Tsurusaki, Fong & Zhang, 2025 introduced by Tan et al. (2025), and eleven species previously assigned to Nelima Roewer, 1910 or Leiobunum are formally transferred to Pseudoliobunum; Paralus Roewer, 1949 was incorrectly considered a junior homonym of Paralus Rafinesque, 1815 (nomen nudum; Mollusca, Gastropoda) by Özdikmen (2006), who proposed the unnecessary replacement name Neoparalus Özdikmen, 2006—this misinterpretation is recognized and corrected here for the first time; Zalmoxis dentata Thorell, 1891 is herein designated as type species of Zalmoxida Roewer, 1912; and Triaenonyx stewartius Hogg, 1910 is herein designated as type species of Metanuncia Roewer, 1915.

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  385. Kury, A.B. (2020a) Incorrect authorship assignments and repeated specific epithets: five cases of the use of homographs by Roewer in harvestmen (Arachnida, Opiliones). Bionomina, 19 (1), 64–76. https://doi.org/10.11646/bionomina.19.1.3
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  392. Kury, A.B. & Cokendolpher, J.C. (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: Kury, A.B., Mendes, A.C., Cardoso, L., Kury, M.S. & Granado, A.de A. (Eds.), WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Self published, Rio de Janeiro, pp. 52–54.
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  394. Kury, A.B., García, A.F. & Ahumada-C., D. (2023) A new genus of Zalmoxoidea from Colombia (Arachnida: Opiliones: Grassatores). The Journal of Arachnology, 51 (1), 37–45. https://doi.org/10.1636/JoA-S-21-070
  395. Kury, A.B., García, A.F. & Medrano, M. (2019) A new genus of Kimulidae; first record of the family from Colombia (Opiliones, Grassatores). Comptes Rendus Biologies, 342, 236–244. https://doi.org/10.1016/j.crvi.2019.07.003
  396. Kury, A.B., Kury, I.S. & Oliveira, A.B.R.de (2024a) Checklists of extant harvestmen (Arachnida: Opiliones) species for all the countries of the world. Zootaxa, 5515 (1), 1–162. https://doi.org/10.11646/zootaxa.5515.1.1
  397. Kury, A.B. & Machado, G. (2009) Notes on Santobius from Vanuatu and Fiji and the status of the eastern Melanesian Ibalonius (Arachnida: Opiliones: Podoctidae). Zoological Studies, 48 (4), 524–538.
  398. Kury, A.B. & Maury, E.A. (1998) A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae). Zoological Journal of the Linnean Society, 123 (2), 143–162. https://doi.org/10.1111/j.1096-3642.1998.tb01297.x
  399. Kury, A.B. & Medrano, M. (2016) Review of terminology for the outline of dorsal scutum in Laniatores (Arachnida, Opiliones). Zootaxa, 4097 (1), 130–134. https://doi.org/10.11646/zootaxa.4097.1.9
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  680. Roewer, C.-F. (1912d) Die Familie der Cosmetiden der Opiliones-Laniatores. Archiv für Naturgeschichte, 78 (10), 1–122, pl. 1–2.
  681. Roewer, C.-F. (1912e) Revision der Opiliones Palpatores (= Opiliones Plagiostethi). II. Teil: Familie der Phalangiidae. (Subfamilien: Sclerosomini, Oligolophini, Phalangiini). Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 20 (1), 1–295, pl. 1–4.
  682. Roewer, C.-F. (1912f) Beitrag zur Kenntnis der Weberknechte Kolumbiens. In: Fuhrmann, O. & Mayor, E. (Eds.), Voyage d’exploration scientifique en Colombie. Mémoires de la Société neuchâteloise des Sciences naturelles, 5, 139–159, pl. 7.
  683. Roewer, C.-F. (1913a) Arachnides. I. Opiliones. In: Résultats scientifiques du voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911–1912). A. Schultz, Paris, pp. 1–22.
  684. Roewer, C.-F. (1913b) Arachnida II: Opiliones. In: Zoological Results of the Abor Expedition, 1911–12. Records of the Indian Museum, 8 (3), 203–207.
  685. Roewer, C.-F. (1913c) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv für Naturgeschichte, 79 (4), 1–256, pl. 1a.
  686. Roewer, C.-F. (1913d) Die Familie der Gonyleptiden der Opiliones - Laniatores. [Part 2]. Archiv für Naturgeschichte, Abteilung A, 79 (5), 257–472.
  687. Roewer, C.-F. (1913e) Opiliones aus N. -Neu-Guinea, gesammelt von P.N. van Kampen und K. Gjellerup in den Jahren 1910 und 1911. Tijdschrift voor Entomologie, 56, 156–164.
  688. Roewer, C.-F. (1914a) Die Opiliones der Sammlung der Herren Drs. Paul u. Fritz Sarasin auf Celebes in den Jahren 1893–1896. Archiv für Naturgeschichte, 79 (10), 70–96.
  689. Roewer, C.-F. (1914b) Arachnida Opiliones. In: Mission du Service Géographique de l’armée pour la mesure d’un Arc de Méridien Équatorial en Amérique du Sud: Vol. Tome 10 (Entomologie, Botanique). Gauthier-Villars et Cie, Paris, pp. 121–141. Issue Fascicule 2 (Opiliones. Diptères. Myriapodes).
  690. Roewer, C.-F. (1914c) Die Familien der Ischyropsalidae und Nemastomatidae der Opiliones=Palpatores. Archiv für Naturgeschichte, 80 (3), 99–169.
  691. Roewer, C.-F. (1914d) Opilioniden von Neu-Caledonien. In: Sarasin, F. & Roux, J. (Eds.), Nova Caledonia, A. Zoologie, 1. C. W. Kreidels, pp. 439–443, pl. 14.
  692. Roewer, C.-F. (1915a) Fünfzehn neue Opilioniden. Archiv für Naturgeschichte, 80 (9), 106–132.
  693. Roewer, C.-F. (1915b) Die Familie der Triaenonychidae der Opiliones - Laniatores. Archiv für Naturgeschichte, 80 (12), 61–168.
  694. Roewer, C.-F. (1915c) 106 neue Opilioniden. Archiv für Naturgeschichte, 81 (3), 1–152.
  695. Roewer, C.-F. (1915d) Neue Opiliones aus dem ungarischen National-Museum in Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 13, 215–223, pl. 4.
  696. Roewer, C.-F. (1916) 7 neue Opilioniden des Zoolog. Museums in Berlin. Archiv für Naturgeschichte, 81 (12), 6–13.
  697. Roewer, C.-F. (1917) 52 neue Opilioniden. Archiv für Naturgeschichte, 82 (2), 90–158.
  698. Roewer, C.-F. (1919) Über Nemastomatiden und ihre Verbreitung. Archiv für Naturgeschichte, 83 (2), 140–160.
  699. Roewer, C.-F. (1920) Australische Opiliones. Results of Dr. E. Mjöberg’s Swedish Scientific Expeditions to Australia 1910-1913. 24. Arkiv för Zoologi, 13 (3–4), 1–11.
  700. Roewer, C.-F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1116 pp.
  701. Roewer, C.-F. (1925) Opilioniden aus Süd-Amerika. Bollettino dei Musei di Zoologia e di Anatomia Comparata della Reale Università di Torino, 40 (34), 1–34, pls. 5–6.
  702. Roewer, C.-F. (1926a) Opiliones vom Mount Maquiling auf Luzon, Philippinen. The Philippine Journal of Science, 29, 545–559.
  703. Roewer, C.-F. (1926b) Opilioniden aus Höhlen des Balkan-Gebirges (Arachn.). Entomologische Mitteilungen, 15, 299–302.
  704. Roewer, C.-F. (1927a) Weitere Weberknechte I. 1. Ergänzung der: “Weberknechte der Erde”, 1923. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen, 26 (2), 261–402, pl. 1.
  705. Roewer, C.-F. (1927b) Brasilianische Opilioniden, gesammelt von Herrn Prof. Bresslau im Jahre 1914. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 40, 331–352.
  706. Roewer, C.-F. (1927c) Ostasiatische Opiliones, von Herrn Prof. F. Silvestri im Jahre 1925 erbeutet. Bollettino del Laboratorio di Zoologia Generale e Agraria della Facoltà Agraria in Portici, 20, 191–210.
  707. Roewer, C.-F. (1928a) Weitere Weberknechte II. (2. Ergänzung der Weberknechte der Erde, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, 26 (3), 527–632, pl. 1.
  708. Roewer, C.-F. (1928b) Opilions nouveaux du Brésil. Bulletin et Annales de la Société Entomologique de Belgique, 68, 123–127.
  709. Roewer, C.-F. (1929a) Contributions to the fauna of Rottnest Island. No. V. Opiliones in the Western Australian Museum. Journal of the Royal Society of Western Australia, 15, 95–98.
  710. Roewer, C.-F. (1929b) On a collection of Indian Palpatores (Phalangiidae) with a revision of the continental genera and species of the sub-family Gagrellinae Thorell. Records of the Indian Museum, 31 (2), 107–159. https://doi.org/10.26515/rzsi/v31/i2/1929/162552
  711. Roewer, C.-F. (1929c) Die Opilioniden der schwedischen Kamtchatka-Expedition 1920–1922. Arkiv för Zoologi, 21 (2), 1–3.
  712. Roewer, C.-F. (1929d) Weitere Weberknechte III. (3. Ergänzung der: “Weberknechte der Erde”, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, 27 (2), 179–284.
  713. Roewer, C.-F. (1929e) Süd-indische Skorpione, Chelonethi und Opilioniden. Revue suisse de Zoologie, 36 (4), 609–639. https://doi.org/10.5962/bhl.part.118944
  714. Roewer, C.-F. (1930) Weitere Weberknechte IV. (4. Ergänzung der Weberknechte der Erde, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, 27 (3), 341–452, pls. 6–7.
  715. Roewer, C.-F. (1931a) Über Triaenonychiden (6. Ergänzung der “Weberknechte der Erde”, 1923). Zeitschrift für wissenschaftliche Zoologie, 138 (1), 137–185.
  716. Roewer, C.-F. (1931b) Weitere Weberknechte V. (5. Ergänzung der “Weberknechte der Erde,” 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, 28 (2–3), 101–164.
  717. Roewer, C.-F. (1931c) Über Opilioniden der Sundainseln. Archiv für Hydrobiologie. Supplement-Band. “Tropische Binnengewässer”, 2, 508–548.
  718. Roewer, C.-F. (1932) Weitere Weberknechte VII (7. Ergänzung der: “Weberknechte der Erde”, 1923) (Cranainae). Archiv für Naturgeschichte (N.F.), 1 (2), 275–350.
  719. Roewer, C.-F. (1933a) Ergebnisse der Österreichischen biologischen Costa-Rica-Expedition 1930. IV. Teil. Opilioniden. Annalen des Naturhistorischen Museums in Wien, 46, 275–295.
  720. Roewer, C.-F. (1933b) Die Opilioniden des Mt. Kinabalu, Nord-Borneo, 13,455 ft. Bulletin of the Raffles Museum, 8, 46–47.
  721. Roewer, C.-F. (1933c) Opiliones aus Perak. Bulletin of the Raffles Museum, 8, 48–51.
  722. Roewer, C.-F. (1934) Änderung des namens einer Opilioniden-Gattung. Zoologischer Anzeiger, 106 (11), 304.
  723. Roewer, C.-F. (1935a) Alte und neue Assamiidae. Weitere Weberknechte VIII (8. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 1 (1), 1–168. 9 pl.
  724. Roewer, C.-F. (1935b) Opiliones. Fünfte Serie, zugleich eine Revision aller bisher bekannten Europäischen Laniatores. Biospeologica. LXII. Archives de Zoologie expérimentale et générale, 78 (1), 1–96.
  725. Roewer, C.-F. (1935c) Südostasiatische Opiliones der Sammlung Fea und Modigliani des Naturhistorischen Museum in Genua. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova, 59, 12–25.
  726. Roewer, C.-F. (1936a) Opiliones aus Malaya. Bulletin of the Raffles Museum, 12, 72–76.
  727. Roewer, C.-F. (1936b) Opilionides von Mauritius. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 1 (3), 335–341, pl. 1.
  728. Roewer, C.-F. (1936c) Zwei sonderbare Pachylinen aus Peru. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 1, 341–343.
  729. Roewer, C.-F. (1938) Über Acrobuninae, Epedaninae und Sarasinicinae. Weitere Weberknechte IX. (9. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 2 (2), 81–169, plates 1–10.
  730. Roewer, C.-F. (1940) Neue Assamiidae und Trogulidae. Weitere Weberknechte X. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 3 (1), 1–31, pl. 1–2.
  731. Roewer, C.-F. (1942a) Opiliones, Pedipalpi und Araneae von Fernando Poo. 21. Beitrag zu den wissenschaftlichen Ergebnissen der Westafrika Expedition Eidmann 1939/40. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 3 (3), 244–258, pl. 19.
  732. Roewer, C.-F. (1942b) Einige neue Arachniden I. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 3, 277–280, pl. 20.
  733. Roewer, C.-F. (1943) Über Gonyleptiden. Weitere Weberknechte (Arachn., Opil.) XI. Senckenbergiana, 26 (1–3), 12–68.
  734. Roewer, C.-F. (1947) Diagnosen neuer Gattungen und Arten der Opiliones Laniatores (Arachn.) aus C.F. Roewer’s Sammlung im Senckenberg-Museum. 1. Cosmetidae. [Weitere Weberknechte XII]. Senckenbergiana, 28 (1/3), 7–57.
  735. Roewer, C.-F. (1949a) Über Phalangodiden I. (Subfam. Phalangodinae, Tricommatinae, Samoinae.) Weitere Weberknechte XIII. Senckenbergiana, 30 (1/3), 11–61.
  736. Roewer, C.-F. (1949b) Einige neue Gattungen der Phalangodidae (Opiliones). Veröffentlichungen aus dem Überseemuseum Bremen, 1, 143–144.
  737. Roewer, C.-F. (1949c) Über Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30 (4/6), 247–289.
  738. Roewer, C.-F. (1950) Opiliones und Solifuga aus Belgisch Congo. Revue de Zoologie et de Botanique Africaines, 44, 30–55.
  739. Roewer, C.-F. (1951a) Einige in Höhlen und Termitenbauten gefundene Assamiidae (Opiliones) aus dem Belgischen Congo-Gebiet. Revue de Zoologie et de Botanique Africaines, 44, 321–327, 3 figs.
  740. Roewer, C.-F. (1951b) Über Nemastomatiden. Weitere Weberknechte XVI. Senckenbergiana, 32 (1/4), 95–153, pls. 1–9.
  741. Roewer, C.-F. (1952a) Solifuga, Opiliones, Pedipalpi und Scorpiones (Arachnoidea). Exploration du Parc National de l’Upemba. Mission G. F. de Witte, 5, 1–36.
  742. Roewer, C.-F. (1952b) Einige Phalangiiden aus dem Vereinigten Staaten von Nord-Amerika. Zoologischer Anzeiger, 149 (11/12), 267–273.
  743. Roewer, C.-F. (1952c) Neotropische Arachnida Arthrogastra, zumeist aus Peru [I]. Senckenbergiana, 33 (1/3), 37–58.
  744. Roewer, C.-F. (1953a) Neotropische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVII). Mitteilungen aus dem Zoologischen Museum in Berlin, 29, 180–265. https://doi.org/10.1002/mmnz.19530290109
  745. Roewer, C.-F. (1953b) Mediterrane Opiliones Palpatores. Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, 33, 201–210.
  746. Roewer, C.-F. (1953c) Opiliones aus Französisch-Westafrika, gesammelt durch Herrn Dr. A. Villiers. Bulletin de l’Institut Français d’Afrique Noire, 15, 610–630.
  747. Roewer, C.-F. (1954a) Spinnentiere aus El Salvador, I. (Arachnoidea: Pedipalpi, Solifuga, Opiliones - Laniatores). Ergebnisse der Forschungsreise A. Zilch 1951 nach El Salvador, 11. Senckenbergiana biologica, 35, 57–73, 10 figs.
  748. Roewer, C.-F. (1954b) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 1. Teil. Senckenbergiana biologica, 35, 181–236.
  749. Roewer, C.-F. (1954c) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 2. Teil. Senckenbergiana biologica, 35 (5/6), 237–292.
  750. Roewer, C.-F. (1954d) Einige neue Opilioniden Laniatoren und Solifugae. Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, 33, 377–384, figs. 1–7.
  751. Roewer, C.-F. (1955a) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 3. Teil. Senckenbergiana biologica, 36, 71–121.
  752. Roewer, C.-F. (1955b) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 4. Teil (Schluss). Senckenbergiana biologica, 36 (3/4), 123–171.
  753. Roewer, C.-F. (1956a) Cavernicole Arachniden aus Sardinien II. Fragmenta Entomologica, 2 (9), 97–104.
  754. Roewer, C.-F. (1956b) Über Phalangiinae (Phalangiidae, Opiliones Palpatores). (Weitere Weberknechte XIX). Senckenbergiana biologica, 37 (3/4), 247–318, pl. 36–43.
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  757. Roewer, C.-F. (1957b) Journey to Northern Ethiopia (Simien), 1952–53: Arachnida, Opiliones. Zoological Journal of the Linnean Society, 43 (290), 93–95. https://doi.org/10.1111/j.1096-3642.1957.tb02509.x
  758. Roewer, C.-F. (1957c) Über Oligolophinae, Caddoinae, Sclerosomatinae, Leiobuninae, Neopilioninae und Leptobuninae (Phalangiidae, Opiliones Palpatores). (Weitere Weberknechte XX). Senckenbergiana biologica, 38 (5/6), 323–358.
  759. Roewer, C.-F. (1959) Neotropische Arachnida Arthrogastra zumeist aus Peru, IV. Senckenbergiana biologica, 40 (1/2), 69–87.
  760. Roewer, C.-F. (1961a) Einige Solifugen und Opilioniden aus der palaearctischen und äthiopischen Region. Senckenbergiana biologica, 42 (5/6), 479–490.
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  763. Roewer, C.-F. (1961d) Opilioniden und Araneen Le Parc National du Niokolo-koba. Fasc. 2. Mémoires de l’Institut Français d’Afrique Noire, 62, 33–81.
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  898. Soares, B.A.M. & Soares, H.E.M. (1949a) Alguns opiliões do sul do Brasil. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 9 (4), 47–60.
  899. Soares, B.A.M. & Soares, H.E.M. (1949b) Monografia dos gêneros de opiliões neotrópicos II. Arquivos de Zoologia do Estado de São Paulo, 7 (2), 149–240. https://doi.org/10.11606/issn.2176-7793.19517149-240
  900. Soares, B.A.M. & Soares, H.E.M. (1954a) Algumas notas sobre opiliões com descrição de novas formas. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 11 (25), 491–507. https://doi.org/10.11606/0031-1049.1954.11p491-507
  901. Soares, B.A.M. & Soares, H.E.M. (1954b) Monografia dos gêneros de opiliões neotrópicos III. Arquivos de Zoologia do Estado de São Paulo, 8 (9), 225–302. https://doi.org/10.11606/issn.2176-7793.19558225-304
  902. Soares, H.E.M. (1945a) Contribuição ao estudo dos opiliões da coleção “Otto Schubart”. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 5 (23), 209–219. https://doi.org/10.11606/0031-1049.1945.5p209-220
  903. Soares, H.E.M. (1945b) Dois novos gêneros e três novas espécies de opiliões brasileiros. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 5 (26), 243–250. https://doi.org/10.11606/0031-1049.1945.5p243-250
  904. Soares, H.E.M. (1946) Contribuição ao estudo dos opiliões do estado do Rio de Janeiro. Revista Brasileira de Biologia, 6 (3), 385–390.
  905. Soares, H.E.M. (1966a) Opiliões da coleção Gofferjè (Opiliones: Gonyleptidae, Phalangodidae). Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 18 (10), 87–102. https://doi.org/10.11606/0031-1049.1966.18p87-102
  906. Soares, H.E.M. (1966b) Novos opiliões da coleção “Otto Schubart” (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 18 (11), 103–115. https://doi.org/10.11606/0031-1049.1966.18p103-115
  907. Soares, H.E.M. (1966c) Opiliões das ilhas dos Búzios e Vitória (Opiliones: Gonyleptidae, Phalangodidae). Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 19 (25), 279–293. https://doi.org/10.11606/0031-1049.1966.19p279-293
  908. Soares, H.E.M. (1968a) Dois novos opiliões do Brasil (Opiliones: Gonyleptidae). Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 21 (14), 137–142. https://doi.org/10.11606/0031-1049.1968.21.p137-142
  909. Soares, H.E.M. (1968b) Contribuição ao estudo dos opiliões do Chile (Opiliones: Gonyleptidae, Triaenonychidae). Papéis Avulsos de Zoologia (São Paulo), 21 (27), 259–272. https://doi.org/10.11606/0031-1049.1968.21.p259-272
  910. Soares, H.E.M. (1970a) Gonyleptids from Poços de Caldas, State of MG, Brazil (Opiliones, Gonyleptidae). Revista Brasileira de Biologia, 30 (2), 211–216.
  911. Soares, H.E.M. (1970b) Novas espécies de opiliões da Região Amazônica (Opiliones, Cosmetidae, Gonyleptidae, Phalangiidae, Stygnidae). Revista Brasileira de Biologia, 30 (3), 323–338.
  912. Soares, H.E.M. (1972) Opera Opiliologica Varia II (Opiliones: Gonyleptidae, Phalangiidae, Phalangodidae). Revista Brasileira de Biologia, 32 (1), 65–74.
  913. Soares, H.E.M. (1977a) Opera Opiliologica Varia VI (Opiliones, Gonyleptidae). Physis. Revista de la Sociedad Argentina de Ciencias Naturales, 36 (92), 251–260.
  914. Soares, H.E.M. (1977b) Opera Opiliologica Varia VII (Opiliones, Gonyleptidae). Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 37 (4), 893–898.
  915. Soares, H.E.M. (1978a) Opera Opiliologica Varia. VIII (Opiliones, Gonyleptidae). Revista Brasileira de Biologia, 38 (1), 73–80.
  916. Soares, H.E.M. (1978b) Opera Opiliologica Varia. XI. Misetropius mendopticus, novo gênero e nova espécie e notas sinonímicas de Stygnidae. Revista Brasileira de Entomologia, 22 (2), 71–76.
  917. Soares, H.E.M. (1978c) Opera Opiliologica Varia. XV. Guatubesia, novo gênero de Gonyleptidae. Revista Brasileira de Entomologia, 22 (2), 99–104.
  918. Soares, H.E.M. (1978d) Opera Opiliologica Varia. XIV. (Opiliones, Phalangodidae). Papéis Avulsos de Zoologia (São Paulo), 32 (12), 141–144. https://doi.org/10.11606/0031-1049.1978.32.p141-144
  919. Soares, H.E.M. (1979a) Opera Opiliologica Varia X (Opiliones, Phalangodidae). Revista Brasileira de Biologia, 39 (1), 161–167.
  920. Soares, H.E.M. (1979b) Novo Caelopyginae do Brasil (Opiliones: Gonyleptidae). Revista Brasileira de Entomologia, 23 (1), 43–46.
  921. Soares, H.E.M. (1979c) Opera Opiliologica Varia XIII (Opiliones, Stygnidae). Revista Brasileira de Biologia, 39 (2), 401–404.
  922. Soares, H.E.M. & Avram, Ş. (1981) Opilionides du Venezuela. Travaux de l’Institut de Spéologie Émile Racovitza, 20, 75–95. https://doi.org/10.59277/tiser
  923. Soares, H.E.M. & Bauab-Vianna, M.J. (1970) Contribución al estudio de los opiliones del Brasil (Opiliones, Gonyleptidae). Physis. Revista de la Sociedad Argentina de Ciencias Naturales, 30 (80), 131–140.
  924. Soares, H.E.M. & Bauab-Vianna, M.J. (1972) Algunas notas sobre opiliones con la descripcion de allotypi y nuevas formas (Opiliones, Gonyleptidae). Physis. Revista de la Sociedad Argentina de Ciencias Naturales, 31 (82), 203–218.
  925. Soares, H.E.M. & Bauab-Vianna, M.J. (1973) Contribuição ao estudo dos Opiliões do Peru (Opiliones: Gonyleptidae). Acta Zoologica Lilloana, 29, 317–342. [“1972”]
  926. Soares, H.E.M. & Soares, B.A.M. (1977) Opera Opiliologica Varia. IX (Opiliones, Gonyleptidae). Physis. Revista de la Sociedad Argentina de Ciencias Naturales, 37 (93), 217–222.
  927. Soares, H.E.M. & Soares, B.A.M. (1979a) Opera Opiliologica Varia XII (Opiliones, Gonyleptidae, Pachylinae). Revista Brasileira de Biologia, 39 (2), 393–399.
  928. Soares, H.E.M. & Soares, B.A.M. (1979b) Opera Opiliologica Varia XVI. Novo gênero de Gonyleptidae e presença de Triaenonychidae no Brasil (Opiliones). Revista Brasileira de Entomologia, 23 (3), 169–173.
  929. Soares, H.E.M. & Soares, B.A.M. (1979c) Opera Opiliologica Varia XVII. (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 23 (4), 257–261.
  930. Soares, H.E.M. & Soares, B.A.M. (1982) Opera Opiliologica Varia XXIII (Opiliones, Gonyleptidae, Gonyleptinae). Revista Brasileira de Entomologia, 26 (1), 15–21.
  931. Soares, H.E.M. & Soares, B.A.M. (1984) Opera opiliologica varia XXV. (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 28 (3), 301–314.
  932. Soares, H.E.M. & Soares, B.A.M. (1985a) Contribution à l’étude des opilions (Opiliones: Cosmetidae, Phalangodidae, Gonyleptidae) du Paraguay. Revue suisse de Zoologie, 92 (1), 3–18. https://doi.org/10.5962/bhl.part.81597
  933. Soares, H.E.M. & Soares, B.A.M. (1985b) Opera Opiliologica Varia XXII. Opiliones Gonyleptidae. Naturalia (São Paulo), 10, 157–200.
  934. Soares, H.E.M. & Soares, B.A.M. (1986) Opera Opiliologica Varia XXVI. (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 30 (1), 87–100.
  935. Soares, H.E.M. & Soares, B.A.M. (1987a) Opera Opiliologica Varia XVIII. (Opiliones, Cosmetidae e Gonyleptidae). Revista Brasileira de Entomologia, 31 (1), 1–11.
  936. Soares, H.E.M. & Soares, B.A.M. (1987b) Opera Opiliologica Varia XIX (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 31 (3), 457–463.
  937. Soares, H.E.M. & Soares, B.A.M. (1987c) Opera Opiliologica Varia XXIV (Opiliones, Gonyleptidae, Pachylinae, Gonyleptinae). Naturalia (São Paulo), 11/12, 53–69.
  938. Soares, H.E.M. & Soares, B.A.M. (1988a) Opera Opiliologica Varia XX. Notas sinonímicas (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 32 (2), 227–232.
  939. Soares, H.E.M. & Soares, B.A.M. (1988b) Opera Opiliologica Varia XXI. (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 32 (2), 233–237.
  940. Soares, H.E.M. & Soares, B.A.M. (1988c) Gonyassamiinae, nova subfamília de Gonyleptidae (Opiliones, Gonyleptoidea, Gonyleptomorphi) do Brasil. Naturalia (São Paulo), 13, 23–27.
  941. Soares, H.E.M., Soares, B.A.M. & Jim, R.de L.S. (1992) Monografia dos gêneros de opiliões neotrópicos IV. (Opiliones, Gonyleptidae, Prostygninae). Revista Brasileira de Entomologia, 36 (1), 1–14.
  942. Sørensen, W.E. (1873) Bidrag til Phalangidernes Morphologi og Systematik samt Beskrivelse af nogle nye, herhen hørende Former. Naturhistorisk Tidsskrift, 8, 489–525, pl. 15. [in Danish]
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  944. Sørensen, W.E. (1884) Opiliones Laniatores (Gonyleptides W. S. Olim) Musei Hauniensis. Naturhistorisk Tidsskrift, 14 (3), 555–646.
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  947. Sørensen, W.E. (1896) Opiliones Laniatores a cl. Dr. Yngwe Sjöstedt in Kamerun (Africa Centrali) collectos. Entomologisk Tidskrift, 17 (2–3), 177–202.
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  980. Suzuki, S. (1963a) A new genus of Gagrellinae (Opiliones) from Japan. Annotationes Zoologicae Japonenses, 36, 97–101.
  981. Suzuki, S. (1963b) Cladolasma parvula gen. et sp. n. (Trogulidae: Opiliones) from Japan. Annotationes Zoologicae Japonenses, 36, 40–44.
  982. Suzuki, S. (1964a) A remarkable new genus of Travuniidae (Phalangida) from Japan. Annotationes Zoologicae Japonenses, 37, 168–173.
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  996. Suzuki, S. (1974) A revision of some harvestmen from Taiwan, with descriptions of two new species (Arachnida, Opiliones, Leiobunidae). Journal of Science of the Hiroshima University, 25, 137–145.
  997. Suzuki, S. (1975) The harvestmen of family Triaenonychidae in Japan and Korea (Travunoidea, Opiliones, Arachnida). Journal of Science of the Hiroshima University, 26, 65–101.
  998. Suzuki, S. (1976a) Two triaenonychid harvestmen from the Northeast, Japan (Triaenonychidae, Opiliones, Arachnida). Journal of Science of the Hiroshima University, 1, 177–185.
  999. Suzuki, S. (1976b) The genus Leiobunum C.L. Koch of Japan and adjacent countries (Leiobunidae, Opiliones, Arachnida). Journal of Science of the Hiroshima University, 1, 187–260.
  1000. Suzuki, S. (1976c) Report on a collection of opilionids from Pasoh Forest Reserve, West Malaysia. Nature and Life in Southeast Asia, 7, 9–38.
  1001. Suzuki, S. (1977) Report on a collection of opilionids from the Philippines. Journal of Science of the Hiroshima University, 27 (1), 1–120.
  1002. Suzuki, S. (1978) Three harvestmen (Arachnida, Opiliones) from the Bonin Islands. Annotationes Zoologicae Japonenses, 51, 179–185.
  1003. Suzuki, S. (1982) Two new genera of Phalangodinae (Opiliones, Phalangodidae) from Eastern Kalimantan, Borneo. Annotationes Zoologicae Japonenses, 55, 100–104.
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