Abstract
We present the first checklist of Asian beetles of the subgenus Copris (Paracopris) Balthasar, 1939 including new data on the distribution. In addition, Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species from Java (Indonesia) is described and Copris (Paracopris) punctulatus Wiedemann, 1823 is redescribed based on specimens deposited in the Muséum National d’Histoire Naturelle (Paris, France), including detailed illustrations of both sexes and male genitalia. At the same time, a neotype is designated for C. (P.) punctulatus as the type specimen was lost.
References
Arrow, G.J. (1931) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp.
Bai, M., Mccullough, E., Song, K.-Q., Liu, W.-G. & Yang, X.-K. (2011) Evolutionary constraints in hind wing shape in Chinese dung beetles (Coleoptera: Scarabaeinae). PLoS ONE, 6 (6), e21600.
https://doi.org/10.1371/journal.pone.0021600
Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R.Università di Firenze. Redia, 25, 1–36.
Balthasar, V. (1958) Eine neue Untergattung und einige neue Arten der Gattung Copris Geoffroy. Acta Entomologica Musei Nationalis Pragae, 32, 471–480.
Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp.
Balthasar, V. (1965) Eine neue Untergattung und neue Arten der familie Scarabaeidae. Acta Entomologica Bohemoslovaca, 62, 14–23.
Bezděk, A. & Hájek, J. (2012) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic (Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini). Acta Entomologica Musei Nationalis Pragae, 52, 297–334.
Bui, V.B., Dumack, K. & Bonkowski, M. (2018) Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species. European Journal of Entomology, 115, 167–191.
https://doi.org/10.14411/eje.2018.016
Chen, K.M., Yang, M.M. & Tsai, J.F. (2002) Dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Lanyu. Collection and Research, 15, 53–62.
Cheung, K., Bai, M., Leung, M., Lee, Y., Chan, K. & Yang, X. (2018) Scarabaeinae (Coleoptera: Scarabaeidae) from Hong Kong. Zoological Systematics, 43 (3), 233–267.
https://doi.org/10.11865/zs.201830
Davis, A.L.V., Frolov, A.V. & Scholtz, C.H. (2008) The African dung beetle genera. Protea Book House, Pretoria, 272 pp.
Fairchild, G.B. (1967) Notes on neotropical Tabanidae (Diptera), VII. The species described by C.R.W. Wiedemann. Pacific Insects, 9 (1), 73–104.
Gupta, D., Chandra, K. & Khan, S. (2014) An updated checklist of scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies, 2 (5), 225–240.
Hanboonsong, Y., Masumoto, K. & Ochi, T. (2003) Dung beetles (Coleoptera, Scarabaeidae) of Thailand. Part 5. Genera Copris and Microcopris (Coprini). Elytra, 31, 103–124.
Hayes, L., Mann, D.J., Monastyrskii, A.L. & Lewis, O.T. (2009) Rapid assessments of tropical dung beetle and butterfly assemblages: contrasting trends along a forest disturbance gradient. Insect Conservation and Diversity, 2, 194–203.
https://doi.org/10.1111/j.1752-4598.2009.00058.x
Horn, W., Kahle, I., Friese, G. & Gaedike, R. (1990) Collectiones Entomologicae. Ein Kompendium über den Verbleib entomologischer Sammlungen der Welt bis 1960. Teil II. L bis Z. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 351 pp. [pp. 223–573]
International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. Fourth edition adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp.
Kabakov, O.N. & Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae of Vietnam and some parts of adjacent countries: South China, Laos, and Thailand. Latvijas Entomologs, 37, 58–96.
Kabakov, O.N. & Shokhin, I.V. (2014) Contribution to the knowledge of the subfamily Scarabaeinae (Coleoptera) from China with nomenclatural notes. Caucasian Entomological Bulletin, 10, 47– 59.
https://doi.org/10.23885/1814-3326-2014-10-1-47-59
Král, D. & Bezděk, A. (2016) Tribe Coprini. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea—Scirtoidea—Dasciloidea—Buprestoidea—Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 167–171.
Löbl, I., Krell, F.-T. & Král, D. (2006) Scarabaeidae: Scarabaeinae: Coprini. In: Löbl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea—Scirtoidea—Dascilloidea—Buprestoidea—Byrrhoidea. Apollo Books, Stenstrup, pp. 151–154.
Mittal, I.C. & Jain, R. (2015) A checklist of Indian dung beetles (Coleoptera: Scarabaeidae). Indian Journal of Entomology, 77 (4), 383–404.
https://doi.org/10.5958/0974-8172.2015.00076.0
Nguyen-Phung, T. (1988) Révision des espèces afrotropicales du genre Copris Müller, 1764 (Coleoptera: Scarabaeidae). IV. Le groupe fidius Olivier (Coleoptera, Scarabaeidae). Revue Française d’Entomologie, Nouvelle Série, 10, 5–15.
Ochi, T. & Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Giornale Italiano di Entomologia, 8, 29–35.
Ochi, T. & Kon, M. (2004) Notes on the coprophagous scarab beetles (Coleoptera, Scarabaeidae) from Southeast Asia (IV). A new horned species of Microcopris from Vietnam and a new subspecies of Copris erratus from Peleng off Sulawesi. Kogane, 5, 25–30.
Ochi, T., Kon, M. & Bai, M. (2009) Three new species of Copris (Coleoptera: Scarabaeidae) from China, with description of a new subgenus. Entomological Review of Japan, 64, 207–216.
Ochi, T., Kon, M. & Kawahara, M. (2008) Four new species of the genus Copris (Coleoptera: Scarabaeidae) from Cambodia and Myanmar. Entomological Review of Japan, 62, 243–253.
Ochi, T., Kon, M. & Kawahara, M. (2019a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469–478.
Ochi, T., Kon, M. & Pham, H.T. (2018) Two new species of Copris (Copris) (Coleoptera: Scarabaeidae) and a new subspecies of Phelotrupes (Sinogeotrupes) strnadi Král, Malý & Schneider (Coleoptera: Geotrupidae) from Vietnam. Giornale Italiano di Entomologia, 15 (63), 159–168.
Ochi, T., Kon, M. & Pham, H.T. (2019b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435–446.
Pont, A.C. (1997) The Muscidae and Fanniidae (Insecta, Diptera) described by C.R.W. Wiedemann. Steenstrupia, 23, 87–122.
Raine, E.H., Gray, C.L., Mann, D.J. & Slade, E.M. (2018) Tropical dung beetle morphological traits predict functional traits and show intraspecific differences across land uses. Ecology and Evolution, 8, 8686–8696.
https://doi.org/10.1002/ece3.4218
Sabu, T.K., Nithya, S. & Vinod, K.V. (2011) Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa, 2830 (1), 29–38.
https://doi.org/10.11646/zootaxa.2830.1.3
Senior, M.J.M., Hamer, K.C., Bottrell, S., Edwards, D.P., Fayle, T.M., Lucey, J.M., Mayhew, P.J., Newton, R., Peh, K.S.H., Sheldon, F.H., Stewart, C., Styring, A.R., Thom, M.D.F., Woodcock, P. & Hill, J.K. (2013) Trait-dependent declines of species following conversion of rain forest to oil palm plantations. Biodiversity and Conservation, 22, 253–268. https://doi.org/10.1007/s10531-012-0419-7
Sewak, R. (2009) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the zoological survey of India, 296, 1–106.
Shahabuddin (2010) Diversity and community structure of dung beetles (Coleoptera: Scarabaeidae) across a habitat disturbance gradient in Lore Lindu National Park, Central Sulawesi. Biodiversitas, 11, 29–33.
https://doi.org/10.13057/biodiv/d110107
Siddiqui, H., Ahmed, Z. & Khatri, I. (2014) Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan journal of zoology, 46 (2), 295–307.
Weidner, H. (1976) Die Entomologischen Sammlungen des Zoologischen Instituts und Zoologischen Museums der Universität Hamburg, IX. Teil, Insecta VI. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut, 73, 87–264.
Wiedemann, C.R.W. (1823) Zweihundert neue Käfer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoologisches Magazin, 2, 1–164.
Zimsen, E. (1954) The insect types of C.R.W. Wiedemann in the Zoological Museum in Copenhagen. Spolia Zoologica Musei Hauniensis, 14, 1–43.