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Type: Article
Published: 2017-08-01
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An annotated checklist of the main representatives of meiobenthos from inland water bodies of Central and Southern Vietnam. I. Roundworms (Nematoda)

Institute for Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Province, 152742, Russia.
Institute for Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Province, 152742, Russia.
Nematoda freshwater Nematoda biodiversity distribution taxonomy population structure and abundance tropics

Abstract

This paper presents an annotated list of nematode taxa found in the course of investigations of meiobenthic communities in various freshwater bodies in central and southern Vietnam. Data on the structure and abundance of populations, ecology, and global and local distribution are given along with taxonomic and other comments. The occurrence and level of taxonomic richness of roundworms in the studied bodies of water and biotopes are discussed briefly. In total, 127 taxa of 10 orders and 38 families of nematodes are listed. The number of recorded genera is 60; 72 representatives of these genera are identified to species level. The listing includes 48 taxa that had not been previously recorded from Vietnamese territory and 17 species originally collected, analyzed, and described as new to science from this country. The results of these investigations and an overview of the previously published data indicate that taxonomic composition and diversity of Nematoda in Vietnamese freshwater habitats remain poorly studied, except for certain areas.

 

References

  1. Ahmad, A. & Ahmad, W. (2002) Studies on the genus Laimydorus Siddiqi, 1969 (Dorylaimida: Dorylaimidae) from India. Journal of Nematode Morphology and Systematics, 4, 91–109.

    Ahmad, W. & Araki, M. (2003) New and known species of the family Dorylaimidae (Nematoda: Dorylaimida) from Japan. International Journal of Nematology, 13, 37–56.

    Ahmad, W. & Jairajpuri, M.S. (1979) First report of the male of Mylonchulus mulveyi Jairajpuri, 1970 (Nematoda: Mononchida). Current Science, 46, 1093–1094.

    Ahmad, W. & Jairajpuri, M.S. (1982) Some new and known species of Dorylaimoidea. Nematologica, 28, 39–61.

    https://doi.org/10.1163/187529282X00501

    Ahmad, W. & Jairajpuri, M.S. (1983) Descriptions of new species of Dorylaimoides and Calolaimus (Dorylaimida) from India. Revue de Nématologie, 6, 65–72.

    Ahmad, W. & Jairajpuri, M.S. (2010) Mononchida: The Predatory Soil Nematodes. Nematology Monographs and Perspectives, 7, 1–300.

    Ahmad, W. & Naz, T. (2012) Four new and six known species of the genus Dorylaimellus Cobb, 1913 (Nematoda: Belondiridae) from India. Journal of Natural History, 46, 2787–2828.

    https://doi.org/10.1080/00222933.2012.724722

    Altherr, E. (1963) Nématodes des sols forestiers subalpins du Val Dischma (Grisons). Bulletin de Societé vaudoise, Lausanne, 68, 333–349.

    Álvarez-Ortega, S., Nguyen, T.A.D., Abolafia, J., Vu, T.T.T. & Peña-Santiago, R. (2015) Three new species of the genus Aporcelaimoides Heyns, 1965 from Vietnam (Nematoda, Dorylaimida, Aporcelaimidae), with an updated taxonomy of the genus. ZooKeys, 516, 1–26.

    https://doi.org/10.3897/zookeys.516.10087

    Álvarez-Ortega, S. & Peña-Santiago, R. (2013) Taxonomy of the genus Aporcelaimellus Heyns, 1965 (Nematoda, Dorylaimida, Aporcelaimidae). Zootaxa, 3669 (3), 243–260.

    https://doi.org/10.11646/zootaxa.3669.3.3

    Álvarez-Ortega, S., Subbotin, S.A. & Peña-Santiago, R. (2013a) Morphological and molecular characterization of Aporcelaimellus simplex (Thorne & Swanger, 1936) Loof & Coomans, 1970 and a new concept for the genus Aporcella Andrássy, 2002 (Dorylaimida, Aporcelaimidae). Nematology, 15, 165–178.

    https://doi.org/10.1163/156854112X651320

    Álvarez-Ortega, S., Subbotin, S.A. & Peña-Santiago, R. (2013b) Morphological and molecular characterisation of Californian species of Metaporcelaimus Lordello, 1965 (Dorylaimida, Aporcelaimidae), with a new concept of the genus. Nematology, 15, 251–278.

    https://doi.org/10.1163/15685411-00002674

    Andrássy, I. (1958) Ergebnisse der zoologischen Aufsammlungen des Ungarischen Naturwissenschaftlichen Museums in Ägypten im Jahre 1957. 2. Nematoden aus ägyptischen Gewässern. Annales Historico-Naturales Musei Nationalis Hungarici, 50, 135–150.

    Andrássy, I. (1964) Neue Nematoden-Arten aus Ungarn, III. Fünf neue Arten. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 5, 9–23.

    Andrássy, I. (1968) Fauna Paraguayensis 2. Nematoden aus den Galeriewäldern des Acaray-Flusses. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 8, 167–315.

    Andrássy, I. (1970) Freilebende Nematoden aus Vietnam. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 10, 5–31.

    Andrássy, I. (1987) The Superfamily Dorylaimoidea (Nematoda)—A Review. Families Thorniidae and Thornenematidae. Acta Zoologica Hungarica, 33, 277–315.

    Andrássy, I. (1988) The superfamily Dorylaimoidea (Nematoda)—a review. Family Dorylaimidae. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 23, 3–63.

    Andrássy, I. (1990) The superfamily Dorylaimoidea (Nematoda)—a review. Family Qudsianematidae, I. Acta Zoologica Hungarica, 36, 163–188.

    Andrássy, I. (1991) The free-living nematode fauna of the Bátorliget nature reserves. In: Mahunka, S. (Ed.), The Bátorliget Nature Reservesafter forty years, Hungarian Natural History Museum, Budapest, 129–197.

    Andrássy, I. (2002a) Free-living nematodes from the Fertö-Hanság National Park, Hungary. In: Mahunka, S. (Eds.), The fauna of Fertö-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 21–97.

    Andrássy, I. (2002b) New genera and species of nematodes from southern Chile. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 34, 5–22.

    Andrássy, I. (2005) Free-living nematodes of Hungary (Nematoda Errantia). I. Pedozoologica Hungarica, No. 3, 1–518.

    Andrássy, I. (2007) Free-living nematodes of Hungary (Nematoda Errantia). II. Pedozoologica Hungarica, No. 4, 1–496.

    Andrássy, I. (2009a) Aporcelinus, a new genus of aporcelaimoid nematodes (Dorylaimida), and its species. International Journal of Nematology, 19, 121–136.

    Andrássy, I. (2009b) Free-living nematodes of Hungary (Nematoda Errantia). III. Pedozoologica Hungarica, No. 5, 1–608.

    Andrássy, I. (2011) A study on Ironus Bastian, 1865 (Nematoda: Еnoplida) with descriptions of two new and two known species. International Journal of Nematology, 21, 27–41.

    Andrássy, I. & Esquivel, A. (2012) Free-living nematodes from nature reserves in Costa Rica Genera Egtitus Thorne, 1967 and Trachypleurosum Andrássy, 1959 (Dorylaimida: Actinolaimidae). Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 43, 3–19.

    Azimi, S., Mahdikhani-Moghadam, E., Rouhani, H. & Memari, H.R. (2016) First report of the nematodes Filenchus orientalis and Hemicriconemoides californianus on faba bean in Iran. Hellenic Plant Protection Journal, 9, 44–50.

    https://doi.org/10.1515/hppj-2016-0005

    Baniyamuddin, M. & Ahmad, W. (2006) New and known species of Dorylaimidae (Nematoda: Dorylaimida) from Arunachal Pradesh, India. Nematology, 8 (4), 491–510.

    https://doi.org/10.1163/156854106778613985

    Baqri, Q.H. & Jairajpuri, M.S. (1968) On six new species of Dorylaimida (Nematoda). Journal of Helminthology, 42, 243–256.

    https://doi.org/10.1017/S0022149X00017831

    Baqri, Q.H. & Jana, A. (1982) Nematodes from West Bengal (India) XIII. Four New Species of Dorylaimidae, with a Key to the Species of Laimydorus Siddiqi, 1969 (Dorylaimoidea). Nematologica, 28, 192–205.

    https://doi.org/10.1163/187529282X00042

    Bert, W. & Borgonie, G. (2006) Order Rhabditida: Suborder Tylenchina. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 648–695.

    https://doi.org/10.1079/9780851990095.0648

    Botha, A. & Heyns, J. (1991) Dorylaimoidea (Nematoda) from rivers in the Kruger National Park. Koedoe, 34, 1–24.

    https://doi.org/10.4102/koedoe.v34i2.420

    Canh, D. & Nguyen, V.T. (2000) Freeliving Nematodes at the Brackish Water Estuary of the Thivai River (Dongnai Province). Tạp chí Sinh Học, 22, 6–9. [in Vietnamese]

    Carbonell, E. & Coomans, A. (1986) The genus Thornenema Andrássy 1959 (Nematoda: Thornenematidae). Nematologica, 32, 125–179.

    https://doi.org/10.1163/187529286x00129

    Coomans, A. & Eyualem, A. (2006) Order Monhysterida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 574–603.

    https://doi.org/10.1079/9780851990095.0574

    Decraemer, W. & Smol, N. (2006) Orders Chromadorida, Desmodorida and Desmoscolecida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 497–573.

    https://doi.org/10.1079/9780851990095.0497

    De Ley, P. & Coomans, A. (1995) Terrestrial nematodes of the Galapagos Archipelago V: Description of Aquatides coboi sp. n., with a key to the genus (Dorylaimida: Nygolaimidae). Russian Journal of Nematology, No. 3 (1), 1–8.

    De Ley, P., Decraemer, W. & Eyualem, A. (2006) Introduction: Summary of Present Knowledge and Research Addressing the Ecology and Taxonomy of Freshwater Nematodes. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 3–30.

    https://doi.org/10.1079/9780851990095.0003

    Eyualem, A. & Coomans, A. (1996) Aquatic nematodes from Ethiopia VII. The family Rhabdolaimidae Chitwood, 1951 sensu Lorenzen, 1981 (Chromadorida: Nematoda) with the description of Udonchus merhatibebi n. sp. Hydrobiologia, 341, 197–214.

    Eyualem, A., Decraemer, W. & De Ley, P. (2008) Global diversity of nematodes (Nematoda) in freshwater. Hydrobiologia, 595, 67–78.

    https://doi.org/10.1007/s10750-007-9005-5

    Eyualem, A., Peng, Y. & Coomans, A. (2001) Thalassomonhystera traesti n. sp., Eumonhystera andrassyi and three Monhystrella species (Monhysteridae: Nematoda) from Li River, China. Hydrobiologia, 462, 185–197.
    https://doi.org/10.1023/A:1013198503784

    Eyualem, A., Traunspurger, W. & Michiels, I.C. (2006) Dynamics of Freshwater Nematodes: Abundance, Biomass and Diversity. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 77–93.

    https://doi.org/10.1079/9780851990095.0077

    Gagarin, V.G. (1992) Free-living Nematodes from the Fresh Waters of USSR. Gidrometeoizdat, St. Petersburg, 152 pp. [in Russian]

    Gagarin, V.G. (1993) Free-living Nematodes from the Fresh Waters of Russia and Adjacent Countries (Order Monhysterida, Areaolaimida, Chromadorida, Enoplida, Mononchida). Gidrometeoizdat, St. Petersburg, 352 pp. [in Russian]

    Gagarin, V.G. (1997) New species of freshwater nematodes of the order Monhysterida from the Novaya Zemlya archipelago (Nematoda). Zoosystematica Rossica, 6, 21–30.

    Gagarin, V.G. (2001) Free-living Nematodes of the Fresh Waters of Russia and Adjacent Countries: Fauna and the Pathways of its Formation, Ecology, Taxonomy, Phylogeny. Nauka, Moscow, 170 pp. [in Russian]

    Gagarin, V.G. & Gusakov, V.A. (2012) Mesodorylaimus dolichurus sp. nov. (Nematoda, Dorylaimida) from freshwater body of Vietnam. Zoosystematica Rossica, 21 (2), 189–192.

    Gagarin, V.G. & Gusakov, V.A. (2013a) Description of Two Species of Free-Living Nematodes (Nematoda) from Fresh Waterbodies of Vietnam. Inland Water Biology, 6, 268–275.

    https://doi.org/10.1134/S199508291304007X

    Gagarin, V.G. & Gusakov, V.A. (2013b) Neotripyla vulgaris gen. n., sp. n. and Semitobrilus andrassyi sp. n. (Nematoda, Triplonchida) from freshwater bodies of Vietnam. Zootaxa, 3716, 565–576.

    https://doi.org/10.11646/zootaxa.3716.4.4

    Gagarin, V.G. & Gusakov, V.A. (2013c) Two New Species of the Family Actinolaimidae (Nematoda) from Water Bodies of Vietnam. Zoologicheskii Zhurnal, 92, 359–365. [in Russian]

    https://doi.org/10.7868/S0044513413010042

    Gagarin, V.G. & Gusakov, V.A. (2013d) Two Species of Dorylaimids (Nematoda) from Waterbodies of Vietnam. Inland Water Biology, 6, 176–183.

    https://doi.org/10.1134/S199508291303005X

    Gagarin, V.G. & Gusakov, V.A. (2014a) Enchodelus minor sp. n. and Mesodorylaimus vietnamicus sp. n. (Nematoda, Dorylaimida) from small freshwater bodies of Vietnam. International Journal of Nematology, 24, 23–30.

    Gagarin, V.G. & Gusakov, V.A. (2014b) Two new species of the genus Laimydorus Siddiqi, 1969 (Nematoda, Dorylaimida) from small freshwater bodies of Vietnam. Amurian Zoological Journal, 6, 117–124.

    Gagarin, V.G. & Gusakov, V.A. (2015a) Enchodelus minor Gagarin & Gusakov, 2014 is a synonym of Dorylaimellus (Axodorylaimellus) parvulus Thorne, 1939. International Journal of Nematology, 25, 215.

    Gagarin, V.G. & Gusakov, V.A. (2015b) Two New Species of Free-Living Nematodes of the Genus Dorylaimoides Thorne, Swanger, 1939 from Fresh Waterbodies of Vietnam. Inland Water Biology, 8, 121–129.

    https://doi.org/10.1134/S1995082915010071

    Gagarin, V.G. & Gusakov, V.A. (2016) Two New Species of Free-Living Nematodes of the Family Dorylaimidae (Nematoda, Dorylaimida) from Small Freshwater Bodies of Vietnam. Inland Water Biology, 9, 242–250.

    https://doi.org/10.1134/S1995082916030056

    Gagarin, V.G. & Nguyen, T.T. (2008a) Free-living nematodes from the Chu River, Northern Vietnam. Inland Water Biology, 1, 14–18.

    https://doi.org/10.1007/s12212-008-1003-4

    Gagarin, V.G. & Nguyen, T.T. (2008b) Free-Living Nematodes from the Red River Delta, Vietnam. Inland Water Biology, 1, 320–325.

    https://doi.org/10.1134/S1995082908040020

    Gagarin, V.G. & Nguyen, V.T. (2003) New species of the genera Chronogaster (Araeolaimida: Chronogasteridae) and Dorylaimellus (Dorylaimida: Belondiridae) from Vietnam (Nematoda). Zoosystematica Rossica, 12, 145–149. [published 20 May 2004]

    Gagarin, V.G. & Nguyen, V.T. (2005a) Free-living nematodes from some fresh water bodies of Northern Vietnam. Biologiya Vnutrennih Vod, No. 1, 18–23. [in Russian]

    Gagarin, V.G. & Nguyen, V.T. (2005b) Three new species of free-living nematodes from freshwater bodies of North Vietnam. International Journal of Nematology, 15, 110–116.

    Gagarin, V.G. & Nguyen, V.T. (2011) Two new species of free-living nematodes from Red River Mouth, Vietnam. International Journal of Nematology, 21, 67–73.

    Gagarin, V.G. & Nguyen, V.T. (2012) Free-Living Nematodes of the Tra Ly River in the Red River Mouth, Vietnam. Inland Water Biology, 5, 11–18.

    https://doi.org/10.1134/S1995082912010038

    Gagarin, V.G., Nguyen, V.T. & Nguyen, D.T. (2003) Three new species of free-living nematodes from freshwater bodies of Vietnam (Nematoda: Araeolaimida). Zoosystematica Rossica, 12, 7–14.

    Giere, O. (2009) Meiobenthology. The microscopic motile fauna of aquatic sediments. Springer-Verlag, Berlin, 527 pp.

    Gusakov, V.A. & Gagarin, V.G. (2012) Meiobenthos Composition and Structure in Highly Mineralized Tributaries of Lake El’ton. Arid Ecosystems, 2, 232–238.

    https://doi.org/10.1134/s2079096112030067

    Gusakov, V.A. & Gagarin, V.G. (2015a) Dorylaimus proximus sp. n. (Nematoda, Dorylaimida) from freshwater bodies of Vietnam. International Journal of Nematology, 25, 117–121.

    Gusakov, V.A. & Gagarin, V.G. (2015b) Two species of free-living nematodes of the order Dorylaimida Pearse, 1942 (Nematoda) from freshwater bodies of Vietnam. Amurian zoological journal, 7, 112–119. [in Russian]

    Gusakov, V.A. & Gagarin, V.G. (2015c) Two Species of Nygolaimidae (Nematoda, Dorylaimida) from Waterbodies of Vietnam. Inland Water Biology, 8, 345–351.

    https://doi.org/10.1134/s1995082915030050

    Gusakov, V.A., Gagarin, V.G. & Semenova, L.M. (2014) Taxonomic composition and structure of meio- and macrozoobenthos of the investigated water bodies. In: Pavlov, D.S. & Zworykin, D.D. (Eds.), Ecology of inland waters of Vietnam. KMK Scientific Press Ltd., Moscow, pp. 163–190. [in Russian]

    Heyns, J. (1968) A monographic study of the nematode families Nygolaimidae and Nygolaimellidae. Entomology Memoirs, No. 19, 1–144.

    Hodda, M. (2007) Phylum Nematoda. In: Zhang, Z.-Q. & Shear, W.A. (Eds.), Linnaeus Tercentenary: Progress in Invertebrate Taxonomy. Zootaxa, 1688, 265–293.

    Hodda, M. (2011) Phylum Nematoda Cobb 1932. In: Zhang, Z.-Q. (Ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 63–95.

    Holovachov, O. & De Ley, P. (2006a) Appendix. Order Mermithida and families incertae sedis (Bastianiidae and Rhabdolaimidae). In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 722–735.

    https://doi.org/10.1079/9780851990095.0722

    Holovachov, O. & De Ley, P. (2006b) Order Plectida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 611–647.

    https://doi.org/10.1079/9780851990095.0611

    Holovachov, O. & Sturhan, D. (2003) Studies on the genus Paraphanolaimus Micoletzky, 1923 (Nematoda: Aphanolaimidae) with description of P. paraguayensis sp. n. Nematology 5, 793–807.

    https://doi.org/10.1163/156854103773040709

    Jacobs, L.J. (1984) The free-living inland aquatic nematodes of Africa—a review. Hydrobiologia, 113, 259–291.

    https://doi.org/10.1007/bf00026614

    Jairajpuri, M.S. (1964) Leptonema thornei n. gen., n. sp. (Nematoda: Dorylaimoidea) from India. Nematologica, 10, 399–402.

    https://doi.org/10.1163/187529264x00402

    Jairajpuri, M.S. & Ahmad, M. (1980) Revised classification of the superfamily Belondiroidea Thorne, 1964 with notes on the systematics of Dorylaimellus Cobb, 1913 (Nematoda: Dorylaimida). Indian Journal of Nematology, 10, 9–22.

    Jairajpuri, M.S. & Khan, W.U. (1982) Predatory Nematodes (Mononchida) with Special Reference to India. Associated Publishing Company, New Delhi, 131 pp.

    Jairajpuri, M.S., Ahmad, M. & Dhanachand, C. (1979) Two New Species of the Genus Thornenema Andrássy, 1959 from Manipur, India. Nematologica, 25, 136–141.

    https://doi.org/10.1163/187529279x00460

    Khan, T.H., Jairajpuri, M.S. & Ahmad, W. (1989) Descriptions of Some New and Known Species of Dorylaim Nematodes. Nematologica, 35, 419–437.

    https://doi.org/10.1163/002825989x00179

    Khan, Z., Ahmad, W. & Jairajpuri, M.S. (1995) Description of new species of dorylaimid nematodes. Afro-Asian Journal of Nematology, 5, 133–140.

    Khan, Z., Ahmad, W. & Rahaman, P.F. (1991) A New Species of the Rare Nematode Genus Trachypleurosum Andrássy, 1959 (Nematoda: Actinolaimidae) from India. Nematologica, 37, 376–381.

    https://doi.org/10.1163/187529291x00385

    Loof, P.A.A. (1973) Freshwater Nematodes from Suriname Collected by J. Van Der Land. Zoologische Verhandelingen, 129, 3–46.

    Loof, P.A.A. (1996) Dichotomous and polytomous identification keys for females of the genera Prodorylaimus Andrássy, 1959 and Laimydorus Siddiqi, 1969 (Nematode: Dorylaimoidea). Russian Journal of Nematology, 4, 7–28.

    Nguyen, T.A.D., Abolafia, J., Bonkowski, M. & Peña-Santiago, R. (2016a) A Third New Species of Aporcelinus Andrássy, 2009 (Dorylaimida, Aporcelaimidae) from Vietnam, with the First SEM Study of a Representative of the Genus. Journal of Nematology, 48, 104–108.

    Nguyen, T.A.D., Abolafia, J., Bonkowski, M. & Peña-Santiago, R. (2016b) Two new species of the genus Aporcelinus Andrássy, 2009 (Nematoda, Dorylaimida, Aporcelaimidae) from Vietnam. Zootaxa, 4103 (6), 550–560.

    https://doi.org/10.11646/zootaxa.4103.6.5

    Nguyen, T.A.D., Vu, T.T.T. & Peña-Santiago, R. (2012) Two new recorded species of the family Leptonchidae (Nematoda: Dorylaimida) for Vietnam’s fauna from Cuc Phuong national park. Tạp chí Sinh Học, 34, 275–282.

    https://doi.org/10.15625/0866-7160/v34n3.2456

    Nguyen, T.A.D., Vu, T.T.T., Bonkowski, M. & Peña-Santiago, R. (2014) New data of three rare belondirid species (Nematoda, Dorylaimida, Belondiridae) from Vietnam, with the first record and description of the male of Oxybelondira paraperplexa Ahmad & Jairajpuri, 1979. Biodiversity Data Journal, 2, 1–15.

    https://doi.org/10.3897/BDJ.2.e1156

    Nguyen, T.T. & Nguyen, V.T. (2005) Comparison on Free-living Nematode Compositions of the Cau, Day, Cam and Nhue Rivers. Tạp chí Sinh Học, 27, 36–42. [in Vietnamese]

    Nguyen, V.T. (2007) Free-living nematodes Monhysterida, Araeolaimida, Chromadorida, Rhabditida, Enoplida, Mononchida, Dorylaimida. In: Dang, N.T. (Ed.), Fauna of Vietnam. Vol. 22. Science and Technics Publishing House, Hanoi, 455 pp. [in Vietnamese]

    Nguyen, V.T. & Gagarin, V.G. (2009) Three Species of Monhysterids (Nematoda, Monhysterida) from Mangrove Forest of the Me Kong River Estuary, Vietnam. Tạp chí Sinh Học, 31, 8–15.

    https://doi.org/10.15625/0866-7160/v31n2.808

    Olia, M., Ahmad, W., Araki, M. & Minaka, N. (2009) Molecular characterization of some species of Mylonchulus (Nematoda: Mononchida) from Japan and comments on the status of Paramylonchulus and Pakmylonchulus. Nematology, 11, 337–342.

    https://doi.org/10.1163/156854109x446935

    Peña-Santiago, R. (2006) Dorylaimida Part I: Superfamilies Belondiroidea, Nygolaimoidea and Tylencholaimoidea. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 326–391.

    https://doi.org/10.1079/9780851990095.0326

    Peña-Santiago, R. & Abolafia, J. (2016) On the identity of Aporcelinus granuliferus (Cobb, 1893) Andrássy, 2009 and its taxonomic consequences. Nematology, 18, 999–1014.

    https://doi.org/10.1163/15685411-00003011

    Peña-Santiago, R. & Ciobanu, M. (2011) The genus Crassolabium Yeates, 1967 (Nematoda, Dorylaimida): a monographic study. Servicio de Publicaciones, Jaén, 195 pp.

    Peña-Santiago, R. & Peralta, M. (1997) The genus Dorylaimoides Thorne & Swanger, 1936 (Nematoda: Dorylaimida). 2. A compendium and key to the species. Fundamental and applied nematology, 20, 253–259.

    Peng, Y., Eyualem, A. & Coomans, A. (2002) A new species of Eumonhystera and three species of Monhystera (Monhysteridae: Nematoda) from Li River, China. Hydrobiologia, 468, 83–93.
    https://doi.org/10.1023/A:1015253123239

    Peralta, M. & Peña-Santiago, R. (1995) Nematodes of the order Dorylaimida from Andalucia Oriental, Spain. The genus Dorylaimoides Thome & Swanger, 1936. 1. Didelphic species. Fundamental and applied nematology, 18, 35–53.

    Peralta, M. & Peña-Santiago, R. (2000) Nematodes of the order Dorylaimida from Andalucia Oriental, Spain. The genus Dorylaimellus Cobb, 1913. Part 1. Journal of Nematode Morphology and Systematics, 2, 121–136.

    Ryss, A.J. (2002) Express technique to prepare collection slides of nematodes. Zoosystematica Rossica, 11, 257–260.

    Schabetsberger, R., Drozdowski, G., Rott, E., Lenzenweger, R., Jersabek, C.D., Fiers, F., Traunspurger, W., Reiff, N., Stoch, F., Kotov, A.A., Martens, K., Schatz, H. & Kaiser, R. (2009) Losing the Bounty? Investigating Species Richness in Isolated Freshwater Ecosystems of Oceania. Pacific Science, 63, 153–179.

    https://doi.org/10.2984/049.063.0201

    Schmidt-Rhaesa, A. (Ed.) (2014) Handbook of zoology: Gastrotricha, Cycloneuralia and Gnathifera. Vol. 2. Nematoda. De Gruyter, Berlin, 759 pp.

    Siddiqi, M.R. (1968) Five New Species of Belondiroidea (Nematoda) from Sibsagar, India, with a Revised Classification of the Superfamily. Proceedings of the Helminthological Society of Washington, 35, 248–258.

    Siddiqi, M.R. (1983) Taxonomy of the subfamily Dorylaimellinae (Nematoda: Dorylaimida). Pakistan Journal of Nematology, 1, 1–38.

    Siddiqi, M.R. & Shah, A.A. (2015) Descriptions of four new species of Amphidorylaimus Andrássy, 1960 (Nematoda: Dorylaimida). International Journal of Nematology, 25, 166–176.

    Siddiqi, M.R., Handoo, Z.A. & Siddiqi, S.F. (2013) A new nematode genus Rugoster (Leptolaimina: Chronogastridae), with descriptions of six new species. International Journal of Nematology, 23, 10–27.

    Smol, N. & Coomans, A. (2006) Order Enoplida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 225–292.

    https://doi.org/10.1079/9780851990095.0225

    Sukul, N.C. (1968) A new species of ironid soil nematode from the Andamans, India, with notes on Oionchus obtusus Cobb. Proceedings of the Zoological Society, 21, 153–156.

    Tahseen, Q., Sultana, R., Khan, R. & Hussain, A. (2012) A New Genus and Species of the Family Rhabdolaimidae (Nematoda), with Descriptions of Two Known Species and Taxonomic Discussion. Journal of Nematology, 44, 302–312.

    Tauheed, U. & Ahmad, W. (2010) Four new species of the genus Mesodorylaimus Andrássy, 1959 (Nematoda: Dorylaimida) from India. Zootaxa, 2642, 19–25.

    Tchesunov, A.V. & Nguyen, V.T. (2010) A description of Anoplostoma nhatrangensis sp. n. from mangrove habitats of Nha Trang, Central Vietnam, with a review of the genus Anoplostoma Bütschli, 1874 (Nematoda: Enoplida). Invertebrate Zoology, 7, 93–105.

    Tchesunov, A.V., Mokievsky, V.O. & Nguyen, V.T. (2010) Three new free-living nematode species (Nematoda, Enoplida) from mangrove habitats of Nha Trang, Central Vietnam. Russian Journal of Nematology, 18, 155–173.

    Timm, R.W. (1964) Nematodes of the superfamily Dorylaimidea from East Pakistan. Proceedings of the Helminthological Society of Washington, 31, 144–153.

    Traunspurger, W., Michiels, I.C. & Eyualem, A. (2006) Composition and Distribution of Free-living Freshwater Nematodes: Global and Local Perspectives. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 46–76.

    https://doi.org/10.1079/9780851990095.0046

    Tsalolikhin, S.Y. (1983) Nematodes of the Families Tobrilidae and Tripylidae of the World Fauna. Nauka, Leningrad, 232 pp. [in Russian]

    Tsalolikhin, S.Y. (1987) Revision of the genus Ironus (Nematoda, Dorylaimida). Zoologicheskii Zhurnal, 66, 661–673. [in Russian]

    Tsalolikhin, S.Y. (1996) Review of the fauna of free-living nematodes from inland waters of Ethiopia. Zoosystematica Rossica, 4, 205–218.

    Tsalolikhin, S.Y. (2009) To the fauna of free-living freshwater nematodes of South-East Asia. Trudy zoologicheskogo instituta RAN, 313, 427–434. [in Russian]

    Tsalolikhin, S.Y. (2015) New species of free-living nematode Monhystera curvicaudata sp. nov. (Nematoda: Monhysterida) from India. Trudy zoologicheskogo instituta RAN, 319, 550–557. [in Russian]

    Vinciguerra, M.T. (2006) Dorylaimida Part II: Superfamily Dorylaimoidea. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 392–467.

    https://doi.org/10.1079/9780851990095.0392

    Vinciguerra, M.T., Orselli, L. & Clausi, M. (2014) One new and two known species of Aporcelinus Andrássy, 2009 and a new species of Coomansinema Ahmad & Jairajpuri, 1989 (Nematoda: Dorylaimida). Nematology, 16, 303–322.

    https://doi.org/10.1163/15685411-00002767

    Vu, T.T.T., Ciobanu, M., Abolafia, J. & Peña-Santiago, R. (2010) Two remarkable new species of the genus Crassolabium Yeates, 1967 from Vietnam (Nematoda: Dorylaimida: Qudsianematidae). Journal of Natural History, 44, 2049–2064.

    https://doi.org/10.1080/00222933.2010.481055

    Wu, J. & Liang, Y. (2000) Inland Free-Living Nematodes in China: A Historical Review of Taxonomic Studies with Descriptions of Aphanonchus orientalis sp. nov. (Chromadorida: Leptolaimidae) and Daptonema limnobia sp. nov. (Monhysterida: Xyalidae). Annales Zoologici, 50, 307–319.

    Yeates, G.W. (1967) Studies on nematodes from dune sands. 6. Dorylaimida. New Zealand Journal of Science, 10, 752–784.

    Zullini, A. (2006) Order Triplonchida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 293–325.

    https://doi.org/10.1079/9780851990095.0293

    Zullini, A. & Peneva, V. (2006) Order Mononchida. In: Eyualem-Abebe, Traunspurger, W. & Andrássy, I. (Eds.), Freshwater nematodes: ecology and taxonomy. CABI Publishing, Wallingford, pp. 468–496.

    https://doi.org/10.1079/9780851990095.0468