Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2003-03-05
Page range: 1–150
Abstract views: 41
PDF downloaded: 4

The Crematogaster (Hymenoptera, Formicidae, Myrmicinae) of Costa Rica

The Evergreen State College, Olympia, Washington 98505, USA
Hymenoptera Crematogaster Costa Rica Formicidae key to species

Abstract

The taxonomy and natural history of the ant genus Crematogaster are reviewed for the Costa Rican fauna. Thirtyone species are known, and a key is provided for these and two additional species from adjacent regions of Panama. Species boundaries are evaluated over their entire range when possible. The taxonomic history of the genus is one of unbridled naming of new species and subspecies, with no synthetic works or keys. Major taxonomic changes are proposed, with the recognition of several polytypic species with very broad ranges and the synonymization of the many names associated with them. Crematogaster pygmaea Forel 1904, suturalis Forel 1912, ornatipilis Wheeler 1918, erici Santschi 1929, and chacoana Santschi 1933 are synonymized under abstinens Forel 1899; centralis Santschi 1932 under acuta (Fabricius 1804); aruga Forel 1913 under arcuata Forel 1899; ludio Forel 1912, armandi Forel 1921, inca Wheeler 1925, and cocciphila Borgmeier 1934 under brasiliensis Mayr 1878; parabiotica Forel 1904 under carinata Mayr 1862; brevispinosa Mayr 1870, minutior Forel 1893, schuppi Forel 1901, recurvispina Forel 1912, sampaioi Forel 1912, striatinota Forel 1912, townsendi Wheeler 1925, and chathamensis Wheeler 1933 under crinosa Mayr 1862; barbouri Weber 1934 under cubaensis Mann 1920; antillana Forel 1893, sculpturata Pergande 1896, kemali Santschi 1923, accola Wheeler 1934, phytoeca Wheeler 1934, panamana Wheeler 1942, and obscura Santschi 1929 under curvispinosa Mayr 1870; descolei Kusnezov 1949 under distans Mayr 1870; projecta Santschi 1925 under erecta Mayr 1866; carbonescens Forel 1913 under evallans Forel 1907; palans Forel 1912, ascendens Wheeler 1925, and dextella Santschi 1929 under limata F. Smith 1858; agnita Wheeler 1934 under obscurata Emery 1895; amazonensis Forel 1905, autruni Mann 1916, and guianensis Crawley 1916 under stollii Forel 1885; surdior Forel 1885, atitlanica Wheeler 1936, and maya Wheeler 1936 under sumichrasti Mayr 1870; tumulifera Forel 1899 and arizonensis Wheeler 1908 under torosa Mayr 1870. The following taxa are raised to species: ampla Forel 1912, brevidentata Forel 1912, chodati Forel 1921, crucis Forel 1912, cubaensis Mann 1920, goeldii Forel 1903, malevolens Santschi 1919, mancocapaci Santschi 1911, moelleri Forel 1912, montana Borgmeier 1939, obscurata Emery 1895, rochai Forel 1903, russata Wheeler 1925, sericea Forel 1912, stigmatica Forel 1911, sub-tonsa Santschi 1925, tenuicula Forel 1904, thalia Forel 1911, uruguayensis Santschi 1912, and vicina Andre 1893. The following new species are described: bryophilia, flavomicrops, flavosensitiva, foliocrypta, jardinero, levior, monteverdensis, raptor, snellingi, sotobosque, and wardi.

References

  1. Adams, E. S. (1994) Territory defense by the ant Azteca trigona: maintenance of an arboreal ant mosaic, Oecologia (Berlin) 97, 203-208.

    André, E. (1893b) Description de quatre espèces nouvelles de fourmis d'Amérique, Revue d'Entomologie (Caen) 12, 148-152.

    Benson, W. W. & Setz, E. Z. F. (1985) On the type localities of ants collected by James Trail in Amazonian Brazil and described by Gustav Mayr, Revista Brasileira de Entomologia 29, 587- 590.

    Bolton, B. (1994) Identification Guide to the Ant Genera of the World, Harvard University Press, Cambridge, Massachusetts, 222 pp.

    Bolton, B. (1995) A New General Catalogue of the Ants of the World, Harvard University Press, Cambridge, Massachusetts, 504 pp.

    Borgmeier, T. (1929) Zur Kenntnis der brasilianischen Ameisen, EOS. Revista Española de Entomología 5, 195-214.

    Borgmeier, T. (1934) Contribuição para o conhecimento da fauna mirmecológica dos cafezais de Paramaribo, Guiana Holandesa (Hym. Formicidae), Archivos do Instituto de Biologia Vegetal (Rio de Janeiro) 1, 93-111.

    Borgmeier, T. (1939) Nova contribuição para o conhecimento das formigas neotropicas (Hym. Formicidae), Revista de Entomologia (Rio de Janeiro) 10, 403-428.

    Buren, W. F. (1959, "1958") A review of the species of Crematogaster, sensu stricto, in North America (Hymenoptera: Formicidae). Part I, Journal of the New York Entomological Society 66, 119-134.

    Buren, W. F. (1968) A review of the species of Crematogaster, sensu stricto, in North America (Hymenoptera, Formicidae). Part II. Descriptions of new species, Journal of the Georgia Entomological Society 3, 91-121.

    Crawley, W. C. (1916) Ants from British Guiana, Annals and Magazine of Natural History (8)17, 366-378.

    Creighton, W. S. (1939) A new subspecies of Crematogaster minutissima with revisionary notes concerning that species (Hymenoptera: Formicidae), Psyche (Cambridge) 46, 137-140.

    Creighton, W. S. (1950) The ants of North America, Bulletin of the Museum of Comparative Zoology 104, 1-585.

    Dalla Torre, K. W. von. (1893) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna), W. Engelmann, Leipzig, 289 pp.

    Davidson, D. W. (1988) Ecological studies of neotropical ant gardens, Ecology 69, 1138-1152. Davidson, D. W., Seidel, J. L. & Epstein, W. W. (1990) Neotropical ant gardens II: bioassays of seed compounds, Journal of Chemical Ecology 16, 2993-3013.

    Emery, C. (1890) Studii sulle formiche della fauna neotropica, Bullettino della Società Entomologica Italiana 22, 38-80.

    Emery, C. (1894) Estudios sobre las hormigas de Costa Rica, Anales del Museo Nacional de Costa Rica 1888-1889, 45-64.

    Emery, C. (1895) Beiträge zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss), Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8, 257-360.

    Emery, C. (1922) Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae [part], Genera Insectorum 174B, 95-206.

    Enzmann, J. (1946) Crematogaster lineolata cerasi, the cherry ant of Asa Fitsch; (with a survey of the American forms of Crematogaster, subgenus Acrocoelia), Journal of the New York Entomological Society 54, 89-97.

    Fabricius, J. C. (1804) Systema Piezatorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus, C. Reichard, Brunswick, xiv + 15-439 + 30 pp.

    Forel, A. (1885, "1884") Études myrmécologiques en 1884 avec une description des organes sensoriels des antennes, Bulletin de la Société Vaudoise des Sciences Naturelles 20, 316-380.

    Forel, A. (1893) Formicides de l'Antille St. Vincent, récoltées par Mons. H. H. Smith, Transactions of the Entomological Society of London 1893, 333-418.

    Forel, A. (1897) Quelques Formicides de l'Antille de Grenada récoltés par M. H. H. Smith, Transactions of the Entomological Society of London 1897, 297-300.

    Forel, A. (1898) La parabiose chez les fourmis, Bulletin de la Société Vaudoise des Sciences Naturelles 34, 380-384.

    Forel, A. (1899) Formicidae [part], Biologia Centrali-Americana, Hymenoptera 3, 81-104.

    Forel, A. (1901a) Einige neue Ameisen aus Südbrasilien, Java, Natal und Mossamedes, Mitteilungen der Schweizerischen Entomologischen Gesellschaft 10, 297-311.

    Forel, A. (1901b) Formiciden des Naturhistorischen Museums zu Hamburg. Neue Calyptomyrmex- , Dacryon-, Podomyrma- und Echinopla-Arten, Mitteilungen aus dem Naturhistorischen Museum in Hamburg 18, 43-82.

    Forel, A. (1903) Mélanges entomologiques, biologiques et autres, Annales de la Société Entomologique de Belgique 47, 249-268.

    Forel, A. (1904a) Miscellanea myrmécologiques, Revue Suisse de Zoologie 12, 1-52.

    Forel, A. (1904b) In und mit Pflanzen lebende Ameisen aus dem Amazonas-Gebiet und aus Peru, gesammelt von Herrn E. Ule, Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 20, 677-707.

    Forel, A. (1907a) Formicides du Musée National Hongrois, Annales Historico-Naturales Musei Nationalis Hungarici 5, 1-42.

    Forel, A. (1907b) Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900, Mitteilungen aus dem Naturhistorischen Museum in Hamburg 24, 1-20.

    Forel, A. (1908a) Fourmis de Costa-Rica récoltées par M. Paul Biolley, Bulletin de la Société Vaudoise des Sciences Naturelles 44, 35-72.

    Forel, A. (1908b) Ameisen aus Sao Paulo (Brasilien), Paraguay etc. gesammelt von Prof. Herm. v. Ihering, Dr. Lutz, Dr. Fiebrig, etc, Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 58, 340-418.

    Forel, A. (1909) Ameisen aus Guatemala usw., Paraguay und Argentinien (Hym.), Deutsche Entomologische Zeitschrift 1909, 239-269.

    Forel, A. (1911a) Ameisen des Herrn Prof. v. Ihering aus Brasilien (Sao Paulo usw.) nebst einigen anderen aus Südamerika und Afrika (Hym.), Deutsche Entomologische Zeitschrift 1911, 285- 312.

    Forel, A. (1911b) Die Ameisen des K. Zoologischen Museums in München, ichte der Mathematischen- Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften zu München 11, 249-303.

    Forel, A. (1912) Formicides néotropiques. Part I, Annales de la Société Entomologique de Belgique 56, 28-49.

    Forel, A. (1913) Fourmis d'Argentine, du Brésil, du Guatémala & de Cuba reçues de M. M. Bruch, Prof. v. Ihering, Mlle Baez, M. Peper et M. Rovereto, Bulletin de la Société Vaudoise des Sciences Naturelles 49, 203-250.

    Forel, A. (1914a) Einige amerikanische Ameisen, Deutsche Entomologische Zeitschrift 1914, 615- 620.

    Forel, A. (1914b) Formicides d'Afrique et d'Amérique nouveaux ou peu connus, Bulletin de la Société Vaudoise des Sciences Naturelles 50, 211-288.

    Forel, A. (1914c) Quelques fourmis de Colombie, Mémoires de la Société Neuchâteloise des Sciences Naturelles 5(2), 9-14.

    Forel, A. (1921, "1920") Fourmis trouvées dans des galles de Cordia et d'Agonandra, etc, Bulletin de la Société Botanique de Genève (2)12, 201-208.

    Forel, A. (1928) The Social World of the Ants Compared with that of Man. Volume 2 [Translated by C. K. Ogden.], G. P. Putnam's Sons, London, xx+444pp.

    Gallardo, A. (1931) Algunas formas sexuales aún no descriptas de las hormigas del género Crematogaster de la República Argentina, Revista de la Sociedad Entomológica Argentina 3, 297- 304.

    Gallardo, A. (1934) Las hormigas de la República Argentina. Subfamilia Mirmicinas, segunda sección Eumyrmicinae, tribu Crematogastrini (Forel), género Crematogaster Lund, Anales del Museo Nacional de Historia Natural de Buenos Aires 38, 1-84.

    Gámez, R. (1991) Biodiversity conservation through facilitation of its sustainable use: Costa Rica's National Biodiversity Institute, Trends in Ecology & Evolution 6, 377-378.

    Heinze, J., Cover, S. P. & Hölldobler, B. (1998, "1995") Neither worker, nor queen: an ant caste specialized in the production of unfertilized eggs, Psyche (Cambridge) 102, 173-185.

    Heinze, J., Foitzik, S., Oberstadt, B., Rüppell, O. & Hölldobler, B. (1999) A female caste specialized for the production of unfertilized eggs in the ant Crematogaster smithi, Naturwissenschaften 86, 93-95.

    Hillis, D. M. (1988) Systematics of the Rana pipiens complex: puzzle and paradigm, Annual Review of Ecology and Systematics 19, 39-63.

    Hölldobler, B., & Wilson, E. O. (1990) The Ants, Harvard University Press, Cambridge, Massachusetts, xii+732pp.

    Holliday, M. (1903) A study of some ergatogynic ants, Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 19, 293-328.

    Kempf, W. W. (1960) Tranopeltoides Wheeler, a synonym of Crematogaster Lund (Hymenoptera, Formicidae), Entomological News 71, 173-175.

    Kempf, W. W. (1968) Miscellaneous studies on Neotropical ants. IV. (Hymenoptera, Formicidae), Studia Entomologica 11, 369-415.

    Kempf, W. W. (1972) Catálogo abreviado das formigas da região Neotropical, Studia Entomologica 15, 3-344.

    Kleinfeldt, S. E. (1978) Ant-gardens: the interaction of codonanthe crassifolia (Gesneriaceae) and Crematogaster longispina (Formicidae), Ecology 59, 449-456.

    Kusnezov, N. (1949) Crematogaster (Neocrema) descolei n. sp. (Hymenoptera, Formicidae), Acta Zoologica Lilloana 8, 587-590.

    Kusnezov, N. (1953) La fauna mirmecológica de Bolivia, Folia Universitaria Cochabamba 6, 211- 229.

    Kutter, H. (1931) Mit Bananen eingeschleppte Ameisen, Mitteilungen der Schweizerischen Entomologischen Gesellschaft 15, 61-64.

    Luederwaldt, H. (1926) Observações biologicas sobre formigas brasileiras especialmente do estado de São Paulo, Revista do Museu Paulista 14, 185-303.

    Longino, J. T., Colwell, R. K. & Coddington, J. A. (2002) The ant fauna of a tropical rainforest: estimating species richness three different ways, Ecology 83, 689-702.

    Maddison, W. P. (1987) Marchena and other jumping spiders with an apparent leg-carapace stridulatory mechanism (Araneae: Salticidae: Heliophaninae and Thiodininae), Bulletin of the British Arachnological Society 7, 101-106.

    Mann, W. M. (1916) The Stanford Expedition to Brazil, 1911, John C. Branner, Director. The ants of Brazil, Bulletin of the Museum of Comparative Zoology 60, 399-490.

    Mann, W. M. (1920) Additions to the ant fauna of the West Indies and Central America, Bulletin of the American Museum of Natural History 42, 403-439.

    Mayr, G. (1862) Myrmecologische Studien, Verhandlungen der Kaiserlich-Königlichen Zoologisch- Botanischen Gesellschaft in Wien 12, 649-776.

    Mayr, G. (1866) Diagnosen neuer und wenig gekannter Formiciden, Verhandlungen der Kaiserlich- Königlichen Zoologisch-Botanischen Gesellschaft in Wien 16, 885-908.

    Mayr, G. (1870a) Formicidae novogranadenses, Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Classe [Klasse]. Abteilung I 61, 370-417.

    Mayr, G. (1870b) Neue Formiciden, Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 20, 939-996.

    Mayr, G. (1878, "1877") Formiciden gesammelt in Brasilien von Professor Trail, Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 27, 867-878.

    Mayr, G. (1887) Südamerikanische Formiciden, Verhandlungen der Kaiserlich-Königlichen Zoologisch- Botanischen Gesellschaft in Wien 37, 511-632.

    Menozzi, C. (1925) Qualche formica nuova od interessante del Deutsch. Ent. Institut di Dahlem (Form.), Entomologische Mitteilungen. Berlin-Dahlem 14, 368-371.

    Menozzi, C. (1929) Formiche di Cuba e delle Isole Canarie raccolte dal Prof. Filippo Silvestri, Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 23, 1-5.

    Mercado, N. K. (1962) Mutualism between Trigona compressa Latr. and Crematogaster stolli Forel (Hymenoptera: Apidae), Journal of the New York Entomological Society 70, 215-217.

    Pergande, T. (1896) Mexican Formicidae, Proceedings of the California Academy of Sciences (2)5, Hölldobler, B., & Wilson, E. O. (1990) The Ants, Harvard University Press, Cambridge, Massachusetts, xii+732pp.

    Holliday, M. (1903) A study of some ergatogynic ants, Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 19, 293-328.

    Kempf, W. W. (1960) Tranopeltoides Wheeler, a synonym of Crematogaster Lund (Hymenoptera, Formicidae), Entomological News 71, 173-175.

    Kempf, W. W. (1968) Miscellaneous studies on Neotropical ants. IV. (Hymenoptera, Formicidae), Studia Entomologica 11, 369-415.

    Kempf, W. W. (1972) Catálogo abreviado das formigas da região Neotropical, Studia Entomologica 15, 3-344.

    Kleinfeldt, S. E. (1978) Ant-gardens: the interaction of codonanthe crassifolia (Gesneriaceae) and Crematogaster longispina (Formicidae), Ecology 59, 449-456.

    Kusnezov, N. (1949) Crematogaster (Neocrema) descolei n. sp. (Hymenoptera, Formicidae), Acta Zoologica Lilloana 8, 587-590.

    Kusnezov, N. (1953) La fauna mirmecológica de Bolivia, Folia Universitaria Cochabamba 6, 211- 229.

    Kutter, H. (1931) Mit Bananen eingeschleppte Ameisen, Mitteilungen der Schweizerischen Entomologischen Gesellschaft 15, 61-64.

    Luederwaldt, H. (1926) Observações biologicas sobre formigas brasileiras especialmente do estado de São Paulo, Revista do Museu Paulista 14, 185-303.

    Longino, J. T., Colwell, R. K. & Coddington, J. A. (2002) The ant fauna of a tropical rainforest: estimating species richness three different ways, Ecology 83, 689-702.

    Maddison, W. P. (1987) Marchena and other jumping spiders with an apparent leg-carapace stridulatory mechanism (Araneae: Salticidae: Heliophaninae and Thiodininae), Bulletin of the British Arachnological Society 7, 101-106.

    Mann, W. M. (1916) The Stanford Expedition to Brazil, 1911, John C. Branner, Director. The ants of Brazil, Bulletin of the Museum of Comparative Zoology 60, 399-490.

    Mann, W. M. (1920) Additions to the ant fauna of the West Indies and Central America, Bulletin of the American Museum of Natural History 42, 403-439.

    Mayr, G. (1862) Myrmecologische Studien, Verhandlungen der Kaiserlich-Königlichen Zoologisch- Botanischen Gesellschaft in Wien 12, 649-776.

    Mayr, G. (1866) Diagnosen neuer und wenig gekannter Formiciden, Verhandlungen der Kaiserlich- Königlichen Zoologisch-Botanischen Gesellschaft in Wien 16, 885-908.

    Mayr, G. (1870a) Formicidae novogranadenses, Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Classe [Klasse]. Abteilung I 61, 370-417.

    Mayr, G. (1870b) Neue Formiciden, Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 20, 939-996.

    Mayr, G. (1878, "1877") Formiciden gesammelt in Brasilien von Professor Trail, Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 27, 867-878.

    Mayr, G. (1887) Südamerikanische Formiciden, Verhandlungen der Kaiserlich-Königlichen Zoologisch- Botanischen Gesellschaft in Wien 37, 511-632.

    Menozzi, C. (1925) Qualche formica nuova od interessante del Deutsch. Ent. Institut di Dahlem (Form.), Entomologische Mitteilungen. Berlin-Dahlem 14, 368-371.

    Menozzi, C. (1929) Formiche di Cuba e delle Isole Canarie raccolte dal Prof. Filippo Silvestri, Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 23, 1-5.

    Mercado, N. K. (1962) Mutualism between Trigona compressa Latr. and Crematogaster stolli Forel (Hymenoptera: Apidae), Journal of the New York Entomological Society 70, 215-217.

    Pergande, T. (1896) Mexican Formicidae, Proceedings of the California Academy of Sciences (2)5, 858-896.

    Roger, J. (1862) Synonymische Bemerkungen. 1. Ueber Formiciden, Berliner Entomologische Zeitschrift 6, 283-297.

    Santschi, F. (1911) Formicides de diverses provenances, Annales de la Société Entomologique de Belgique 55, 278-287.

    Santschi, F. (1912) Quelques fourmis de l'Amérique australe, Revue Suisse de Zoologie 20, 519- 534.

    Santschi, F. (1913) Hyménoptères. Formicides. In: André, E. et al. (Ed) Mission du service géographique de l'armée pour la mesure d'un arc de méridien équatorial en Amérique du Sud. Tome 10. Fasc. 1. Insectes, Gauthier-Villars, Paris, 33-43.

    Santschi, F. (1918) Sous-genres et synoymies [sic] de Cremastogaster (Hym. Formic.), Bulletin de la Société Entomologique de France 1918, 182-185.

    Santschi, F. (1919) Nouveaux formicides de la République Argentine, Anales de la Sociedad Cientifica Argentina 87, 37-57.

    Santschi, F. (1922) Description de nouvelles fourmis de l'Argentine et pays limitrophes, Anales de la Sociedad Cientifica Argentina 94, 241-262.

    Santschi, F. (1923) Solenopsis et autres fourmis néotropicales, Revue Suisse de Zoologie 30, 245- 273.

    Santschi, F. (1925) Nouveaux Formicides brésiliens et autres, Bulletin et Annales de la Société Entomologique de Belgique 65, 221-247.

    Santschi, F. (1929a) Nouvelles fourmis de la République Argentine et du Brésil, Anales de la Sociedad Cientifica Argentina 107, 273-316.

    Santschi, F. (1929b) Mélange myrmécologique, Wiener Entomologische Zeitung 46, 84-93.

    Santschi, F. (1932) Quelques fourmis inédites de l'Amérique centrale et Cuba, Revista de Entomologia (Rio de Janeiro) 2, 410-414.

    Santschi, F. (1933) Fourmis de la République Argentine en particulier du territoire de Misiones, Anales de la Sociedad Cientifica Argentina 116, 105-124.

    Santschi, F. (1939) Résultats scientifiques des croisières du navire-école belge, "Mercator". XIV. Formicidae s. lt, Mémoires du Musée Royal d'Histoire Naturelle de Belgique (2)15, 159-167.

    Seidel, J. L., Epstein, W. W. & Davidson, D. W.. (1990) Neotropical ant gardens I: chemical constituents, Journal of Chemical Ecology 16, 1791-1816.

    Shepard, M., & Gibson, F. (1972) Spider-ant symbiosis: Cotinusa spp. (Araneida: Salticidae) and Tapinoma melanocephalum (Hymenoptera: Formicidae), Canadian Entomologist 104, 1951- 1954.

    Skwarra, E. (1934a) Ökologie der Lebensgemeinschaften mexikanischer Ameisenpflanzen, Zeitschrift für Morphologie und Ökologie der Tiere 29, 306-373.

    Skwarra, E. (1934b) Ökologische Studien über Ameisen und Ameisenpflanzen in Mexiko, published by author (printer R. Leupold), Königsberg, 153 pp.

    Smith, F. (1857) Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace [part], Journal and Proceedings of the Linnean Society of London. Zoology 2, 42-88.

    Smith, F. (1858) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part VI. Formicidae, British Museum, London, 216 pp.

    Soulié, J. (1961, "1960") Des considérations écologiques peuvent-elles apporter une contribution à la connaisance du cycle biologique des colonies de Cremastogaster (Hymenoptera - Formicoidea)? Insectes Sociaux 7, 283-295.

    Weber, N. A. (1934) Notes on neotropical ants, including the descriptions of new forms, Revista de Entomologia (Rio de Janeiro) 4, 22-59.

    Weber, N. A. (1938) New ants from stomachs of Bufo marinus L. and Typhlops reticulatus (L), Annals of the Entomological Society of America 31, 207-210.

    Wheeler, D. E. (1986) Ectatomma tuberculatum foraging biology and association with Crematogaster (Hymenoptera Formicidae), Annals of the Entomological Society of America 79, 300- 303.

    Wheeler, G. C., & Wheeler, J. (1952) The ant larvae of the myrmicine tribe Crematogastrini, Journal of the Washington Academy of Sciences 42, 248-262.

    Wheeler, G. C., & Wheeler, J. (1960) Supplementary studies on the larvae of the Myrmicinae (Hymenoptera: Formicidae), Proceedings of the Entomological Society of Washington 62, 1- 32.

    Wheeler, W. M. (1906) The habits of the tent-building ant (Cremastogaster lineolata Say), Bulletin of the American Museum of Natural History 22, 1-18.

    Wheeler, W. M. (1908a) The ants of Jamaica, Bulletin of the American Museum of Natural History 24, 159-163.

    Wheeler, W. M. (1908b) The ants of Texas, New Mexico and Arizona. (Part I.), Bulletin of the American Museum of Natural History 24, 399-485.

    Wheeler, W. M. (1910) Ants: their structure, development and behavior, Columbia University Press, New York, xxv+663pp.

    Wheeler, W. M. (1912) Notes on a mistletoe ant, Journal of the New York Entomological Society 20, 130-133.

    Wheeler, W. M. (1917) Jamaican ants collected by Prof. C. T. Brues, Bulletin of the Museum of Comparative Zoology 61, 457-471.

    Wheeler, W. M. (1918) Ants collected in British Guiana by Mr. C. William Beebe, Journal of the New York Entomological Society 26, 23-28.

    Wheeler, W. M. (1921a) A new case of parabiosis and the "ant gardens" of British Guiana, Ecology 2, 89-103.

    Wheeler, W. M. (1921b) The Tachigalia ants, Zoologica (N. Y.) 3, 137-168.

    Wheeler, W. M. (1922a) Neotropical ants of the genera Carebara, Tranopelta and Tranopeltoides, new genus, American Museum Novitates 48, 1-14.

    Wheeler, W. M. (1922b) Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VII. Keys to the genera and subgenera of ants, Bulletin of the American Museum of Natural History 45, 631-710.

    Wheeler, W. M. (1923) Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn Dr. A. Roman in Amazonas 1914-1915. 7. Formicidae, Arkiv för Zoologi 15(7), 1-6.

    Wheeler, W. M. (1925) Neotropical ants in the collections of the Royal Museum of Stockholm, Arkiv för Zoologi 17A(8), 1-55.

    Wheeler, W. M. (1933) The Templeton Crocker Expedition of the California Academy of Sciences, 1932. No. 6. Formicidae of the Templeton Crocker Expedition, Proceedings of the California Academy of Sciences (4)21, 57-64.

    Wheeler, W. M. (1934) Neotropical ants collected by Dr. Elisabeth Skwarra and others, Bulletin of the Museum of Comparative Zoology 77, 157-240.

    Wheeler, W. M. (1936) A singular Crematogaster from Guatemala, Psyche (Cambridge) 43, 40-48.

    Wheeler, W. M. (1942) Studies of Neotropical ant-plants and their ants, Bulletin of the Museum of Comparative Zoology 90, 1-262.

    Wheeler, W. M. & Darlington, P. J., Jr. (1930) Ant-tree notes from Rio Frio, Colombia, Psyche (Cambridge) 37, 107-117.