Abstract
The taxonomy of the Southeast Asian mite harvestman family Stylocellidae is updated in light of new molecular and mor-phometric phylogenies, examinations of type specimens, and a new species from Thailand. A new genus, Giribetia gen.nov., is erected, and Fangensis insulanus Schwendinger & Giribet, 2005, recombined in it as Giribetia insulana newcomb. All species in the genus Stylocellus have been recombined in Miopsalis and Leptopsalis, except for the type species,S. sumatranus Westwood, 1874, and a new species, S. lornei, sp. nov., described here. The new recombinations of formerStylocellus species are as follows: Leptopsalis pangrango (Shear, 1993), new comb., Leptopsalis sedgwicki (Shear, 1979),new comb., Leptopsalis laevichelis (Roewer, 1942), new comb., Miopsalis globosa (Schwendinger & Giribet, 2004), newcomb., Miopsalis kinabalu (Shear, 1993), new comb., Miopsalis leakeyi (Shear, 1993), new comb., Miopsalis mulu(Shear, 1993), new comb., and Miopsalis pocockii (Hansen & Sørensen, 1904), new comb. “Stylocellus” spinifronsRoewer, 1942 is now designated as nomen dubium, as the sole type specimen has been found to be a juvenile. Two newsubfamilies are proposed, each with two genera: Fangensinae subfam. nov., containing Fangensis and Giribetia, and Lep-topsalinae subfam. nov., containing Leptopsalis and Miopsalis. The subfamily sensu strictu Stylocellinae contains the remaining two genera: Stylocellus and MeghalayaReferences
Bastawade, D.B. (1992) First report of an arachnid order Cyphophthalmi (da) from India in Arunachal Pradesh. Journal of the Bombay Natural History Society, 89, 268–269.
Clouse, R., General, D., Diesmos, A. & Giribet, G. (2011) An old lineage of Cyphophthalmi (Opiliones) discovered on Mindanao highlights the need for biogeographical research in the Philippines. Journal of Arachnology, 39, 147–153.
Clouse, R.M., de Bivort, B.L. & Giribet, G. (2009) A phylogenetic analysis for the South-east Asian mite harvestman family Stylocellidae (Opiliones : Cyphophthalmi)—a combined analysis using morphometric and molecular data. Invertebrate Systematics, 23, 515–529.
Clouse, R.M. & Giribet, G. (2007) Across Lydekker's Line — first report of mite harvestmen (Opiliones : Cyphophthalmi : Stylocellidae) from New Guinea. Invertebrate Systematics, 21, 207–227.
Clouse, R.M. & Giribet, G. (2010) When Thailand was an island — the phylogeny and biogeography of mite harvestmen (Opiliones, Cyphophthalmi, Stylocellidae) in Southeast Asia. Journal of Biogeography, 37, 1114–1130.
Clouse, R. & Giribet, G. (in press) On the Cyphophthalmi (Arachnida, Opiliones) types from the Museo Civico di Storia Naturale "Giacomo Doria". Bulletin of the Museum of Comparative Zoology.
Colwell, R.K. (2004-2011) Biota 2: The Biodiversity Database Manager. Sunderland: Sinauer Associates.
de Bivort, B.L., Clouse, R.M. & Giribet, G. (2010) A morphometrics-based phylogeny of the temperate Gondwanan mite harvestmen (Opiliones, Cyphophthalmi, Pettalidae). Journal of Zoological Systematics and Evolutionary Research, 48, 294–309.
Giribet, G. (2000) Catalogue of the Cyphophthalmi of the world (Arachnida, Opiliones). Revista Iberica de Aracnologia, 2, 49–76.
Giribet, G. (2002) Stylocellus ramblae, a new Stylocellid (Opiliones, Cyphophthalmi) from Singapore, with a discussion of the family Stylocellidae. Journal of Arachnology, 30, 1–9.
Giribet, G. & Boyer, S.L. (2002) A cladistic analysis of the cyphophthalmid genera (Opiliones, Cyphophthalmi). Journal of Arachnology, 30, 110–128.
Giribet, G., Sharma, P., Benavides, L., Boyer, S., Clouse, R., de Bivort, B.L., Dimitrov, D., Kawauchi, G.Y., Murienne, J. & Schwendinger, P. (2012) Evolutionary and biogeographic history of the harvestman suborder Cyphophthalmi (Arachnida, Opiliones)—an ancient and global group of arachnids. Biological Journal of the Linnean Society, 105, 92–130.
Giribet, G., Sharma, P.P. & Bastawade, D.B. (2007) A new genus and species of Cyphophthalmi (Arachnida : Opiliones) from the north-eastern states of India. Zoological Journal of the Linnean Society, 151, 663–670.
Hansen, H.J. & Sørensen, W. (1904) On two orders of Arachnida: Opiliones, especially the suborder Cyphophthalmi, and Ricinulei, namely the family Cryptostemmatoidae. Cambridge University Press, Cambridge.
Karsch, F. (1884) Ueber die Arachniden-Familie der Sironoiden und über einen neuen Siro aus Asturien. Berliner entomologische Zeitschrift, 28, 143-146.
Lohman, D.J., de Bruyn, M., Page, T., von Rintelen, K., Hall, R., Ng, P.K.L., Shih, H.-T., Carvalho, G.R. & von Rintelen, T. (2011) Biogeography of the Indo-Australian Archipelago. Annual Review of Ecology, Evolution and Systematics, 42, 205–226.
Murphree, C.S. (1988) Morphology of the dorsal integument of ten Opilionid Species (Arachnida, Opiliones). Journal of Arachnology, 16, 237–252.
Pocock, R.I. (1897) Descriptions of some new Oriental Opiliones recently received by the British Museum. Annals and Magazine of Natural History, Series 6, 19, 283-292.
Rambla, M. (1991) A new Stylocellus from some caves of Borneo, Malaysia (Opiliones, Cyphophthalmi, Stylocellidae). Memoires de Biospeologie, 18, 227–232.
Rambla, M. (1994) Un nouveau Cyphophthalme du sud-est asiatique, Fangensis leclerci n. gen. n. sp. (Opiliones, Sironidae). Memoires de Biospeologie, 21, 109–114.
Roewer, C.F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1116 pp.
Roewer, C.F. (1927) Weitere Weberknechte I. (I. Ergänzung der: "Weberknechte der Erde," 1923.). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen, Bremen, 26, 261-402.
Roewer, C.F. (1942) Einige neue Arachniden I. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, Bremen, 3, 277–280.
Schwendinger, P.J. & Giribet, G. (2005) The systematics of the south-east Asian genus Fangensis Rambla (Opiliones : Cyphophthalmi : Stylocellidae). Invertebrate Systematics, 19, 297–323.
Schwendinger, P.J., Giribet, G. & Steiner, H. (2004) A remarkable new cave-dwelling Stylocellus (Opiliones, Cyphophthalmi) from peninsular Malaysia, with a discussion on taxonomic characters in the family Stylocellidae. Journal of Natural History, 38(11), 1421–1435.
Shear, W.A. (1979) Slylocellus sedgwicki n. sp., from Penang Island, Malaysia (Opiliones, Cyphophthalmida, Stylocellidae). Bulletin of the British Arachnological Society, 4, 356–360.
Shear, W.A. (1980) A review of the Cyphophthalmi of the United States and Mexico, with a proposed reclassification of the suborder (Arachnida, Opiliones). American Museum Novitates, 2705, 1–34.
Shear, W.A. (1993) New species in the opilionid genus Stylocellus from Malaysia, Indonesia and the Philippines (Opiliones, Cyphophthalmi, Stylocellidae). Bulletin of the British Arachnological Society, 9, 174–188.
Simon, E. (1879) Les Arachnides de France, Tome Septième, Contenant les ordres des Chernetes, Scorpiones et Opiliones. Émile Martinet, Paris, 332 pp.
Stelbrink, B., Albrecht, C., Hall, R. & von Rintelen, T. (2012) The biogeography of Sulawesi revisited: is there evidence for a vicariant origin of taxa on Wallace’s “anomalous island”? Evolution, 66, 2252–2271
Sundevall, C.J. (1833) Conspectus Arachnidum. C.F. Berling, Londini Gothorum [Lund (Sweden)], 39 pp.
Thorell, T. (1882) Descrizione di alcuni Aracnidi inferiori dell' Arcipelago Malese. Annali del Museo Civico di Storia Naturale di Genova, 18, 21–69.
Thorell, T. (1890a) Aracnidi di Nias e di Sumatra raccolti nel 1886 dal Sig. E. Mondigliani. Annali del Museo Civico di Storia Naturale di Genova, 2a, 10, 5–106.
Thorell, T. (1890b) Aracnidi di Pinang raccolti nel 1889 dai Sig.ri L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 2a, 10, 269–383.
Westwood, J.O. (1874) Thesaurus Entomologicus Oxoniensis; or, illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F.W. Hope, M.A., D.C.L., F.R.S., &c. with forty plates from drawings by the author. Clarendon Press, Oxford.