Abstract
Eleven Afrotropical species of Bengalia Robineau-Desvoidy, ten in the Bengalia peuhi species-group and one reassignedto the Bengalia spinifemorata species-group, are revised. The male genitalia of all, and the ovipositor of six species, areillustrated by means of digital colour photography. A key to males is provided. Two species, i.e., Bengalia minor Malloch,1927 and Bengalia peuhi Villeneuve, 1914, are assigned to the Bengalia peuhi subgroup of the Bengalia peuhi species-group. Five species, i.e., Bengalia depressa Walker, 1858, Bengalia floccosa Wulp, 1885, Bengalia gaillardi Surcouf &Guyon, 1912, Bengalia roubaudi Rickenbach, Hamon & Mouchet, 1960 and Bengalia tibiaria Villeneuve, 1926 areassigned to the Bengalia floccosa subgroup of the Bengalia peuhi species-group. Three species, i.e., Bengalia africanoidessp. nov., Bengalia aliena Malloch, 1927 and Bengalia wyatti (Lehrer, 2005) comb. nov., have been left as species incertaesedis in the Bengalia peuhi species-group. The eleventh species, Bengalia bantuphalla (Lehrer, 2005) comb. nov., is re-assigned to the Bengalia spinifemorata species-group. Bengalia africanoides sp. nov. has only 3 post dc setae, a featureunique for a species of Bengalia. A neotype is designated for Calliphora floccosa Wulp, 1885 (now in Bengalia), to fixthe interpretation of the name in accordance with current usage. Lectotypes are designated for Bengalia bekilyana Séguy,1935, Bengalia depressa, Bengalia gaillardi, Bengalia peuhi and Bengalia unicalcarata Villeneuve, 1913 to fix theinterpretation of the names. Bengalia africana Malloch, 1927 is established as a junior synonym of Bengalia depressa,syn. nov. Bengalia cuthbertsoni Zumpt, 1956 is established as a junior synonym of Bengalia tibiaria, syn. nov.Shakaniella sakinehae Lehrer, 2011 is established as a junior synonym of Shakaniella wyatti (now in Bengalia), syn. nov.Tsunamia yourubana Lehrer, 2005 is established as a junior synonym of Bengalia aliena, syn. nov. Ochromyia petersianaLoew, 1852 is removed from its current position as a doubtful senior synonym of Bengalia depressa and transferred to thegenus Thoracites Brauer & Bergenstamm, 1891 as Thoracites petersiana, comb. nov. in the Rhiniidae. This name is proposed as a senior synonym of Thoracites neglectus Zumpt, 1972, syn. nov.References
Andrews, P. (2008) Advanced Photoshop Elements 6 for Digital Photographers. Focal Press (Elsevier). viii + 413 pp.
Bezzi, M. (1892) Di alcuni Ditteri raccolti nel paese dei Somali dall’ingegnere L. Bricchetti-Robecchi. Annali del Museo civico di Storia Naturale di Genova, 32, 181–196.
Bezzi, M. (1908) Ditteri Eritrei raccolti dal Dott. Andreini e dal Prof. Tellini. Parte Seconda (1) Diptera Cyclorrhapha. Bullettino della Società Entomologica Italiana, 39 [1907], 3–199.
Bezzi, M. (1911) Miodarii superiori raccolti dal signor C. W. Howard nell’Africa australe orientale. Bollettino del Laboratorio di Zoologia Generale e Agraria della R. Scuola Superiore d’Agricoltura in Portici, 6 [1912], 45–104.
Bezzi, M. (1913) Einige Bemerkungen über die Dipterengattungen Auchmeromyia und Bengalia. Entomologische Mitteilungen, 2, 70–78.
Bezzi, M. (1914) Ditteri raccolti dal Prof. F. Silvestri durante il suo viaggio in Africa del 1912–13. Bollettino del Laboratorio di Zoologia Generale e Agraria della R. Scuola Superiore d’Agricoltura in Portici, 8 [1913–1914], 279–308.
Bigot, J.M.F. (1888) Diptères nouveaux ou peu connus. Muscidi (J.B.). Bulletin de la Société Zoologique de France, 12 [1887], 581–617.
Brauer, F. (1897) Beiträge zur Kenntniss der Muscaria schizometopa und Beschreibung von zwei Hypoderma-Arten. I. Bemerkungen zu den Originalexemplaren der von Bigot und Macquart beschriebenen Muscaria schizometopa aus der Sammlung des Herrn G. H. Verrall. Sitzungsberichte der kaiserliche Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, Abt. 1, 106, 329–377.
Brauer, F. (1898) Beiträge zur Kenntniss der Muscaria schizometopa. [Part.] I. Bemerkungen zu den Originalexemplaren der von Bigot, Macquart und Robineau-Desvoidy beschriebenen Muscaria schizometopa aus der Sammlung des Herrn G. H. Verrall. Zweite Folge. Sitzungsberichte der kaiserliche Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, Abt. 1, 107, 493–546.
Brauer, F. (1899) Beiträge zur Kenntniss der Muscaria schizometopa. Bemerkungen zu den Originalexemplaren der von Bigot, Macquart und Robineau-Desvoidy beschriebenen Muscaria schizometopa aus der Sammlung des Herrn G. H. Verrall. Dritte Folge. Sitzungsberichte der kaiserliche Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, Abt. 1, 108, 495–529.
Brauer, F. & Bergenstamm, J.E. von (1891) Die Zweiflügler des kaiserlichen Museums zu Wien. V. Vorarbeiten zu einer Monographie der Muscaria Schizometopa (exclusive Anthomyidae). Pars II. F. Tempsky, Wien. 142 pp.
Crosskey, R.W. (1974) The British Tachinidae of Walker and Stephens (Diptera). Bulletin of the British Museum (Natural History), Entomology, 30, 269–308.
Dawah, H.A. & Abdullah, M.A. (2009) The Calliphoridae (Diptera: Cyclorrhapha) of south-western Saudi Arabia. Fauna of Arabia, 24, 359–371.
Deeming, J.C. (1996) The Calliphoridae (Diptera: Cyclorrhapha) of Oman. Fauna of Saudi Arabia, 15, 264–279.
Deeming, J.C. (2008) Order Diptera, family Calliphoridae. Arthropod fauna of the UAE, 1, 724–731.
Duponchel, P. (1842) Bengalie. In: Orbigny, C.V.D. d’ (ed.), Dictionnaire Universel d’Histoire Naturelle. Tome deuxième. C. Renard, Paris, p. 542.
Note. For bibliographic details on the Dictionnaire and the type designations therein, see Evenhuis & Thompson (1990).
Evenhuis, N.L. & Thompson, F.C. (1990) Type designations of genus-group names of Diptera given in d’Orbigny’s Dictionnaire Universel d’Histoire Naturelle. Bishop Museum Occasional Papers, 30, 226–258.
Fabricius, J.C. (1805) Systema antliatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae [= Brunswick]. xiv + 15–372 + [1 (Errata)] + 30 pp.
Feng, Y., Chen, H.-w. & Xue, W.-q. (1998) Calliphoridae. In: Xue W.-q. & Chao C.-m. (eds), Flies of China. Vol. 2. Liaoning Science and Technology Press, Shenyang [1996], pp. 1366–1517. [In Chinese, with English summaries of new species.]
International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. Fourth edition adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London. xxix + 306 pp.
James, M.T. (1964) Diptera from Nepal. The blow flies (Diptera: Calliphoridae). Bulletin of the British Museum (Natural History), Entomology, 15, 171–179.
James, M.T. (1977) Family Calliphoridae. In: Delfinado, M.D. & Hardy, D.E. (eds), A catalog of the Diptera of the Oriental Region. Volume III Suborder Cyclorrhapha (excluding Division Aschiza). University Press of Hawaii, Honolulu, pp. 526–556.
Karsch, F. (1888) Bericht über die durch Herrn Lieutenant Dr. Carl Wilhelm Schmidt in Ost-Afrika gesammelten und von der zoologischen Abtheilung des Königlichen Museums für Naturkunde in Berlin erworbenen Dipteren. Berliner entomologische Zeitschrift, 31 [1887], 367–382.
Kurahashi, H. (2001) Four new species of the blow fly genus Thoracites Brauer & Bergenstamm (Diptera: Calliphoridae: Rhiniinae), with a key to all species. Cimbebasia, 17, 143–161.
Kurahashi, H. & Kirk-Spriggs, A.H. (2006) The Calliphoridae of Namibia (Diptera: Oestroidea). Zootaxa, 1322, 1–131.
Lehrer, A.Z. (2005) Bengaliidae du Monde (Insecta, Diptera). Pensoft Series Faunistica 50, Pensoft Publishers, Sofia, Moscow. 192 pp.
Lehrer, A.Z. (2006) Contributions taxonomiques et zoogéographique sur la famille des Bengaliidae (Diptera). Bulletin de la Société Entomologique de Mulhouse, 62, 1–11.
Lehrer, A.Z. (2011a) Deux nouveaux Bengaliides de la faune de Malawi (Diptera, Bengalidae). Fragmenta dipterologica, 28, 7–10. [Issue 28 was received as a pdf file from the author on 2 March 2011.]
Lehrer, A.Z. (2011b) Monstruosités diptérologiques de Knut Rognes avec l’assentiment des superviseurs zootaxistes (Diptera, Bengaliidae). Fragmenta dipterologica, 30, 1–14. [This issue was received as a pdf file from the author on 29 July 2011.]
Loew, H. (1852) [Hr. Peters legte Diagnosen und Abbildungen der von ihm in Mossambique neu entdeckten Dipteren vor, welche von Hrn. Professor Loew bearbeitet worden sind.] Bericht über die zur Bekanntmachung geeigneten Verhandlungen der Königlichen Preussische Akademie der Wissenschaften zu Berlin, 1852, 658–661.
Loew, H. (1862) Diptera Zweiflügler. In: Peters, W.C.H. (ed.), Naturwissentschaftliche Reise nach Mossambique auf Befehl seiner Majestät des Königs Friedrich Wilhelm IV in den Jahren 1842 bis 1848 ausgeführt. Zoologie, 5 (Insecten und Myriapoden), pp. 1–34. [xxi + 566 pp. + 35 pls.]
Malloch, J.R. (1927) LV.—Exotic Muscaridæ (Diptera).—XX. Annals and Magazine of Natural History, (9) 20, 385–424.
Peris, S.V. (1952) La subfamilia Rhiniinae (Dipt. Calliphoridae). Anales de la Estacion Experimental de Aula Dei, 3, 1–224.
Pont, A.C. (1980) 90. Family Calliphoridae. In: Crosskey, R.W. (ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 779–800.
Rickenbach, A. (1967) Calliphoridae de République centrafricaine avec la description d’une nouvelle espèce: Isomyia yvoreyi n. sp. [Dipt. Cyclorrhapha]. Bulletin de la Société Entomologique de France, 72, 45–52.
Rickenbach, A., Hamon, J. & Mouchet, J. (1960) Description de deux Calliphoridae nouveaux de la région éthiopienne: Bengalia roubaudi n. sp. et Tricyclea colasbelcouri n. sp. (Diptera; Cyclorhapha). Bulletin de la Société de Pathologie exotique et de ses Filiales, 53, 155–158.
Rickenbach, A., Hamon, J. & Ovazza, M. (1962) Calliphoridae de Haute Volta et de Côte d’Ivoire avec quelques notes sur ceux du Mali, du Sénégal, de la Guinée et du Dahomey [Dipt. Cyclorhapha]. Bulletin de la Société Entomologique de France, 67, 132–141.
Robineau-Desvoidy, J.-B. (1830) Essai sur les Myodaires. Mémoires présentés par divers savants à l’Académie Royale des Sciences de l’Institut de France, Sciences Mathémathiques et Physiques, 2, 1–813.
Rognes, K. (1991) Blowflies (Diptera, Calliphoridae) of Fennoscandia and Denmark. Fauna entomologica Scandinavica, 24, 1–272.
Rognes, K. (1997) The Calliphoridae (blowflies) (Diptera: Oestroidea) are not a monophyletic group. Cladistics, 13, 27–66.
Rognes, K. (2006) Bengalomania – A review of Andy Z. Lehrer’s book on Bengalia Robineau-Desvoidy, 1830 and related works (Diptera, Calliphoridae). Studia dipterologica, 12 [2005], 443–471.
Rognes, K. (2009) Revision of the Oriental species of the Bengalia peuhi species-group (Diptera, Calliphoridae). Zootaxa, 2251, 1–76.
Rognes, K. (2011a) Revision of the Bengalia spinifemorata species-group (Diptera, Calliphoridae). Zootaxa, 2835, 1–29.
Rognes, K. (2011b) A review of the monophyly and composition of the Bengaliinae with the description of a new genus and species, and new evidence for the presence of Melanomyinae in the Afrotropical Region (Diptera, Calliphoridae). Zootaxa, 2964, 1–60.
Séguy, E. (1933). Contributions à l’étude de la faune du Mozambique. Voyage de M. P. Lesne (1928–1929). 13.e Note.- Diptères (2.e partie). Memórias e estudos do Museu Zoológico da Universidade de Coimbra. Série 1., No.67 (1933), 5–80.
Séguy, E. (1935) Caracteres particuliers des calliphorines. Encyclopédie entomologique. Série B. Mémoires et Notes. II. Diptera, 8, 121–150.
Séguy, E. (1946) Calliphorides d’Extreme-Orient. Encyclopédie entomologique. Série B. Mémoires et Notes. II. Diptera, 10, 81–90.
Séguy, E. (1958) Les Calliphores africaines du Muséum (Diptères). Bulletin de l’Institut français d’Afrique noire, Série A, 20,142–192.
Senior-White, R.A., Aubertin, D. & Smart, J. (1940) The fauna of British India, including the remainder of the Oriental Region. Diptera. Vol. VI. Calliphoridæ. Taylor & Francis, London, xiii + 288 pp.
Surcouf, J.M.R. [1920] Révision des Muscidæ testaceæ. Nouvelles Archives du Muséum d’Histoire Naturelle de Paris, (5) 6 (‘1914’, ‘1919’), 27–124.
Note. For details about the publication date of this work, see the summary by Rognes (2009: 73).
Surcouf, J.M.R. & Guyon, L. (1912) Nouvelles espèces de Calliphorinæ de l’Afrique occidentale. Bulletin du Muséum National d’Histoire Naturelle, 18, 418–428.
Vaughan, M. (1991) Curing their Ills. Colonial Power and African Illness. Stanford University Press, Stanford, California. 224 pp.
Villeneuve, J. (1913a) Sur le genre Bengalia Rob.-Desv. [Dipt.]. Bulletin de la Société entomologique de France, 1913 (6), 151–153. [Séance du 26 mars 1913.] [Published 9 April 1913.]
Villeneuve, J. (1913b) Description d’une espèce nouvelle du genre Bengalia Rob.-Desv. [Dipt.]. Bulletin de la Société Entomologique de France, 1913 (6), 153–154. [Séance du 26 mars 1913.] [Published 9 April 1913.]
Villeneuve, J. (1913c) Note rectificative [Dipt.]. Bulletin de la Société Entomologique de France, 1913 (14), 348. [Séance du 23 juillet 1913.] [Published 13 August 1913.]
Villeneuve, J. (1914) Liste d’espèces africaines du genre Bengalia Rob.-Desv. [Dipt.]. Bulletin de la Société Entomologique de France, 1914 (8), 253–255. [Séance du 22 avril 1914.] [Published 13 May 1914.]
Villeneuve, J. (1926) Myodaires supérieurs de l’Afrique nouvaux ou peu connus. Revue Zoologique Africaine, 14, 64–69.
Walker, F. (1858) Characters of undescribed Diptera in the collection of W. W. Saunders, Esq., F.R.S., &c. [part]. Transactions of the Entomological Society of London, N. Ser. 4, 190–235 [or: … London, (2) 4, 190–235].
Wiedemann, C.R.W. (1819) Beschreibung neuer Zweiflügler aus Ostindien und Afrika. Zoologisches Magazin (Kiel), 1 (3), 1–39.
Wulp, F.M. van der (1885) Quelques Diptères exotiques. Annales de la Société Entomologique de Belgique, 28 [1884], CCLXXXVIII–CCXCVII.
Zumpt, F. (1956) Calliphoridae (Diptera Cyclorrhapha) Part I: Calliphorini and Chrysomyiini. Exploration du Parc National Albert Mission G. F. de Witte (1933–1935), 87, 1–200.
Zumpt, F. (1958) Calliphoridae (Diptera Cyclorrapha) Part II: Rhiniini. Exploration du Parc National Albert Mission G. F. de Witte (1933–1935), 92, 1–207.
Zumpt, F. (1962a) The Calliphoridae of the Madagascan Region (Diptera). Part I. Calliphorinae. Verhandlungen der Naturforschenden Gesellschaft in Basel, 73, 41–100.
Zumpt, F. (1962b) Notes on Calliphoridae from the Ethiopian Region, with descriptions of three new species (Diptera). Journal of the Entomological Society of Southern Africa, 25, 236–254.
Zumpt, F. (1972) Thoracites B.B. in the Ethiopian Region, with descriptions of two new species (Diptera: Sarcophagidae). Journal of the New York Entomological Society, 80, 48–53.
Zumpt, F. & Stimie, M. (1965) Notes on Calliphoridae of the Ethiopian Region, with descriptions of eight new species (Diptera). Annals of the Natal Museum, 18, 3–19.